Eleocharis bolanderi |
Eleocharis cylindrica |
|
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Bolander's spikerush |
cylinder spikerush |
|
Habit | Plants perennial, densely tufted; rhizomes caudexlike, mostly hidden by culms and roots, short, 1.5–3 mm thick, hard, cortex persistent, internodes very short, scales not evident. | Plants perennial; rhizomes evident, 1 mm thick, firm, cortex unknown, internodes unknown, scales fugacious, 5 mm, membranous. |
Culms | subterete, often with to 6 prominent ridges when dry, sulcate, 10–30 cm × 0.3–0.5 mm, firm to rigid, spongy. |
terete, often with a few blunt ridges when dry, 20–50 cm × 0.2–0.7 mm, soft, spongy. |
Leaves | distal leaf sheaths persistent, not splitting, proximally brown, red, or stramineous, distally stramineous, green or reddish, papery, apex sometimes reddish, obtuse, rarely callose, tooth absent. |
distal leaf sheaths persistent, not splitting, proximally red or stramineous, distally green, membranous, apex usually reddish, subtruncate to obtuse, not callose, tooth present, to 0.4 mm. |
Spikelets | ovoid, 3–8 × 2–3 mm, apex acute to obtuse; proximal scale amplexicaulous, entire; subproximal scale with flower; floral scales spreading in fruit, 8–30, 4–5 per mm of rachilla, dark brown to blackish, midrib regions often stramineous or greenish, ovate to lanceolate, 2–3 × 1.5 mm, apex entire, acute, often carinate in distal part of spikelet. |
narrowly lanceoloid, narrowly ovoid, or narrowly cylindric, 4–10 × 1–1.5 mm, apex narrowly acute; proximal scale amplexicaulous or clasping 3/4 of culm, entire; subproximal scale empty or with flower; floral scales spreading in fruit, 20–80, 5–6 per mm of rachilla, orange-brown, midrib regions stramineous, narrowly ovate-lanceolate, 1.5–2 × 0.8 mm, entire, apex acute, not carinate. |
Flowers | perianth bristles 3–6, whitish to stramineous, stout to slender, often unequal, from rudimentary to 1/2 of achene length; stamens 3; anthers dark yellow to brown, 0.9–1.4 mm; styles 3-fid. |
perianth bristles 3–6, stramineous, stout, unequal, rudimentary to less than 1/2 achene length; stamens 3; anthers stramineous, 0.7–1.4 mm; styles 3-fid. |
Achenes | falling with scales, stramineous, rarely dark brown, ovoid to obpyriform, slightly to greatly compressed-trigonous, rarely thickly lenticular, angles prominent or abaxial angle obscure, 0.9–1.2 × 0.65–0.8 mm, apex narrowly to broadly truncate, neck short, often compressed more than body, at 20–30X finely rugulose with more than 20 horizontal ridges in a vertical series or reticulate or cancellate. |
falling with scales, stramineous, ellipsoid, compressed-trigonous, angles prominent (keeled), 0.6–0.65 × 0.4–0.45 mm, base narrowed, neck long, smooth at 40X. |
Tubercles | whitish to brown, pyramidal, lower than wide, often 3-lobed as viewed from the top, 0.1–0.3 × 0.4–0.65 mm. |
brown, pyramidal, as high as wide to greatly depressed, 0.1–0.25 × 0.15–0.2 mm. |
Eleocharis bolanderi |
Eleocharis cylindrica |
|
Phenology | Fruiting late spring–summer. | Fruiting spring–summer. |
Habitat | Fresh, often summer-dry meadows, springs, seeps, stream margins | Ephemeral pools in old rock quarry |
Elevation | 1000–3400 m (3300–11200 ft) | 300 m (1000 ft) |
Distribution |
CA; CO; ID; NV; OR; UT
|
TX; South America (Argentina, Paraguay) |
Discussion | Eleocharis bolanderi is clearly distinct from E. montevidensis, from which it differs in its dense, tufted habit with short, caudexlike rhizomes, its leaf sheaths without a tooth, its achene and tubercle shapes, and its acute floral scales. Specimens of Eleocharis bolanderi without rhizomes or achenes are easily confused with E. decumbens, which often may be distinguished by culms 0.5–2 mm wide, and spikelets with scales sometimes more than 3 mm long. The tubercles of E. bolanderi are usually poorly developed and much lower than wide; in E. decumbens they are usually well developed and about as high as wide. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Eleocharis cylindrica is synonymous with E. spegazzinii Barros, which was described from temperate South America (E. R. Guaglianone and O. Ueno 1990). It is very uncommon in North America, where it has been confirmed from Aransas, Burnet, and Cameron counties in Texas; reports from Lubbock and Presidio counties have not been confirmed. The report from New Mexico cannot be confirmed because the specimens lack achenes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23. | FNA vol. 23, p. 81. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis |
Sibling taxa | ||
Synonyms | E. montevidensis var. bolanderi | E. texana |
Name authority | A. Gray: Proc. Amer. Acad. Arts 7: 392. (1868) | Buckley: Proc. Acad. Nat. Sci. Philadelphia 14: 10. (1863) |
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