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Bolander's spikerush

two-tone spikerush

Habit Plants perennial, densely tufted; rhizomes caudexlike, mostly hidden by culms and roots, short, 1.5–3 mm thick, hard, cortex persistent, internodes very short, scales not evident. Plants probably perennial, tufted, not stoloniferous; rhizomes present, ± horizontal, 0.2–0.4 mm thick, firm, longer internodes 3–7 mm, scales translucent or not.
Culms

subterete, often with to 6 prominent ridges when dry, sulcate, 10–30 cm × 0.3–0.5 mm, firm to rigid, spongy.

often ascending, often spotted or streaked red-brown, cross section quadrangular or broadly elliptic, often sulcate, 1.5–8(–25) cm × 0.1–0.3 mm, soft.

Leaves

distal leaf sheaths persistent, not splitting, proximally brown, red, or stramineous, distally stramineous, green or reddish, papery, apex sometimes reddish, obtuse, rarely callose, tooth absent.

distal leaf sheaths persistent or fugaceous, closely sheathing, not conspicuously wrinkled, colorless or pale brown, sometimes spotted red-brown, delicately membranous, apex narrowly acute.

Spikelets

ovoid, 3–8 × 2–3 mm, apex acute to obtuse;

proximal scale amplexicaulous, entire;

subproximal scale with flower;

floral scales spreading in fruit, 8–30, 4–5 per mm of rachilla, dark brown to blackish, midrib regions often stramineous or greenish, ovate to lanceolate, 2–3 × 1.5 mm, apex entire, acute, often carinate in distal part of spikelet.

basal spikelets absent; never proliferous, ovoid or ellipsoid, laterally compressed or terete, 1.5–3.7 × 0.8–3 mm, apex acute;

proximal scale empty, persistent, amplexicaulous, similar to floral scales;

subproximal scale with a flower;

floral scales distichous (spirodistichous), 8–22, 6–10 per mm of rachilla, colorless or pale brown, mottled red-brown to purple, ovate, (1–)2 × 1 mm, membranous, midribs green and red-brown, prominent, apex rounded to obtuse.

Flowers

perianth bristles 3–6, whitish to stramineous, stout to slender, often unequal, from rudimentary to 1/2 of achene length;

stamens 3;

anthers dark yellow to brown, 0.9–1.4 mm;

styles 3-fid.

perianth bristles present or absent, (1–)4, colorless to white, vestigial to shorter than achene, 0–0.55 mm;

spinules not evident at 45X;

stamens (1–)3;

anthers 0.35–0.45 mm;

styles 2-fid (or 3-fid).

Achenes

falling with scales, stramineous, rarely dark brown, ovoid to obpyriform, slightly to greatly compressed-trigonous, rarely thickly lenticular, angles prominent or abaxial angle obscure, 0.9–1.2 × 0.65–0.8 mm, apex narrowly to broadly truncate, neck short, often compressed more than body, at 20–30X finely rugulose with more than 20 horizontal ridges in a vertical series or reticulate or cancellate.

whitish to pale brown, obovoid, biconvex, rarely trigonous, angles prominent, 0.5–0.8 × 0.4–0.6 mm, apex slightly constricted proximal to tubercle, smooth.

Tubercles

whitish to brown, pyramidal, lower than wide, often 3-lobed as viewed from the top, 0.1–0.3 × 0.4–0.65 mm.

sessile, green or red-brown, pyramidal, birettaform or umbonate, 0.15–0.25 × 0.3–0.4 mm.

Eleocharis bolanderi

Eleocharis bicolor

Phenology Fruiting late spring–summer. Fruiting summer–late fall.
Habitat Fresh, often summer-dry meadows, springs, seeps, stream margins Moist banks, freshwater, damp pine barrens
Elevation 1000–3400 m (3300–11200 ft) 0–50 m (0–200 ft)
Distribution
from FNA
CA; CO; ID; NV; OR; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
FL; GA; MS; West Indies; Central America (Nicaragua)
Discussion

Eleocharis bolanderi is clearly distinct from E. montevidensis, from which it differs in its dense, tufted habit with short, caudexlike rhizomes, its leaf sheaths without a tooth, its achene and tubercle shapes, and its acute floral scales.

Specimens of Eleocharis bolanderi without rhizomes or achenes are easily confused with E. decumbens, which often may be distinguished by culms 0.5–2 mm wide, and spikelets with scales sometimes more than 3 mm long. The tubercles of E. bolanderi are usually poorly developed and much lower than wide; in E. decumbens they are usually well developed and about as high as wide.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

The culms of Eleocharis bicolor sometimes have complete transverse septa which are prominent when plants are aquatic.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23. FNA vol. 23, p. 96.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms E. montevidensis var. bolanderi
Name authority A. Gray: Proc. Amer. Acad. Arts 7: 392. (1868) Chapman: Fl. South. U.S., 517. (1860)
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