Eleocharis bolanderi |
Eleocharis bernardina |
|
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Bolander's spikerush |
fewflower spikerush |
|
Habit | Plants perennial, densely tufted; rhizomes caudexlike, mostly hidden by culms and roots, short, 1.5–3 mm thick, hard, cortex persistent, internodes very short, scales not evident. | Plants perennial; rhizomes 0.5–1 mm thick, scales persistent, 5–8 mm, membranous, not fibrous; resting buds absent (non-resting buds on rhizome apex ellipsoid, 10 × 2–5 mm); caudices present, hard, 1–2 mm thick. |
Culms | subterete, often with to 6 prominent ridges when dry, sulcate, 10–30 cm × 0.3–0.5 mm, firm to rigid, spongy. |
markedly arched, not spirally twisted, not contracted near spikelet, when dry usually with several blunt to acute ridges and sulcate, subterete to slightly compressed, to 2 times as wide as thick, 5–15 cm × 0.5–1 mm, firm to hard; culm tufts not proximally bulbous. |
Leaves | distal leaf sheaths persistent, not splitting, proximally brown, red, or stramineous, distally stramineous, green or reddish, papery, apex sometimes reddish, obtuse, rarely callose, tooth absent. |
distal leaf sheaths brown to reddish proximally, stramineous to reddish distally, apex sometimes reddish, membranous to papery. |
Spikelets | ovoid, 3–8 × 2–3 mm, apex acute to obtuse; proximal scale amplexicaulous, entire; subproximal scale with flower; floral scales spreading in fruit, 8–30, 4–5 per mm of rachilla, dark brown to blackish, midrib regions often stramineous or greenish, ovate to lanceolate, 2–3 × 1.5 mm, apex entire, acute, often carinate in distal part of spikelet. |
3–6 × 2–4 mm; proximal scale empty, 2.5–4 mm, 1/2 or more as long as spikelet; floral scales 4–9 per spikelet, 3.5–5 × 2 mm. |
Flowers | perianth bristles 3–6, whitish to stramineous, stout to slender, often unequal, from rudimentary to 1/2 of achene length; stamens 3; anthers dark yellow to brown, 0.9–1.4 mm; styles 3-fid. |
perianth bristles 4–7, very unequal, some shorter than 1/2 of achene, stout and smooth, others equaling or exceeding tubercle, very slender and densely spinulose; anthers 1–1.5 mm. |
Achenes | falling with scales, stramineous, rarely dark brown, ovoid to obpyriform, slightly to greatly compressed-trigonous, rarely thickly lenticular, angles prominent or abaxial angle obscure, 0.9–1.2 × 0.65–0.8 mm, apex narrowly to broadly truncate, neck short, often compressed more than body, at 20–30X finely rugulose with more than 20 horizontal ridges in a vertical series or reticulate or cancellate. |
medium brown, equilaterally trigonous to compressed trigonous or some biconvex, 1.5–2.1 × 0.9–1.2 mm, beak 0.2–0.3 × 0.3–0.4 mm. |
Tubercles | whitish to brown, pyramidal, lower than wide, often 3-lobed as viewed from the top, 0.1–0.3 × 0.4–0.65 mm. |
0.25–0.5 × 0.15–0.2 mm. |
Eleocharis bolanderi |
Eleocharis bernardina |
|
Phenology | Fruiting late spring–summer. | Fruiting summer. |
Habitat | Fresh, often summer-dry meadows, springs, seeps, stream margins | Wet meadows in conifer forests |
Elevation | 1000–3400 m (3300–11200 ft) | 2100–2700 m (6900–8900 ft) |
Distribution |
CA; CO; ID; NV; OR; UT
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CA |
Discussion | Eleocharis bolanderi is clearly distinct from E. montevidensis, from which it differs in its dense, tufted habit with short, caudexlike rhizomes, its leaf sheaths without a tooth, its achene and tubercle shapes, and its acute floral scales. Specimens of Eleocharis bolanderi without rhizomes or achenes are easily confused with E. decumbens, which often may be distinguished by culms 0.5–2 mm wide, and spikelets with scales sometimes more than 3 mm long. The tubercles of E. bolanderi are usually poorly developed and much lower than wide; in E. decumbens they are usually well developed and about as high as wide. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Although usually included in Eleocharis quinqueflora, E. bernardina is clearly more like E. suksdorfiana. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23. | FNA vol. 23. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis | Cyperaceae > Eleocharis > subg. Zinserlingia |
Sibling taxa | ||
Synonyms | E. montevidensis var. bolanderi | Scirpus bernardinus, E. pauciflora var. bernardina, E. quinqueflora var. bernardina |
Name authority | A. Gray: Proc. Amer. Acad. Arts 7: 392. (1868) | (Munz & I. M. Johnston) Munz: Man. S. Calif. Bot., 68, 597. (1935) |
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