Eleocharis bolanderi |
Eleocharis bella |
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Bolander's spikerush |
beautiful spikerush, delicate spikerush, pretty spikerush |
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Habit | Plants perennial, densely tufted; rhizomes caudexlike, mostly hidden by culms and roots, short, 1.5–3 mm thick, hard, cortex persistent, internodes very short, scales not evident. | Plants annual, rarely perennial, usually densely tufted; rhizomes rarely evident, 0.2–0.3 mm thick, internodes 1–5 mm, scales not evident. |
Culms | subterete, often with to 6 prominent ridges when dry, sulcate, 10–30 cm × 0.3–0.5 mm, firm to rigid, spongy. |
often ascending or spreading, 4-angled or sometimes terete, sometimes sulcate, 1–7 cm × 0.2–0.3 mm, soft to firm. |
Leaves | distal leaf sheaths persistent, not splitting, proximally brown, red, or stramineous, distally stramineous, green or reddish, papery, apex sometimes reddish, obtuse, rarely callose, tooth absent. |
sheaths stramineous, distal sheaths often splitting abaxially, slightly inflated distally, oblique, apex acute. |
Spikelets | ovoid, 3–8 × 2–3 mm, apex acute to obtuse; proximal scale amplexicaulous, entire; subproximal scale with flower; floral scales spreading in fruit, 8–30, 4–5 per mm of rachilla, dark brown to blackish, midrib regions often stramineous or greenish, ovate to lanceolate, 2–3 × 1.5 mm, apex entire, acute, often carinate in distal part of spikelet. |
ovoid, 1.5–4 × 0.8–2 mm, apex acute; floral scales 4–15, 8 per mm of rachilla, colorless or reddish brown, midrib region green, ovate-lanceolate, not folded lengthwise, 1–1.5 × 0.5–0.7 mm, mibrib obscure to somewhat keeled, apex narrowly acute to acuminate, slightly recurved. |
Flowers | perianth bristles 3–6, whitish to stramineous, stout to slender, often unequal, from rudimentary to 1/2 of achene length; stamens 3; anthers dark yellow to brown, 0.9–1.4 mm; styles 3-fid. |
perianth bristles absent; anthers 0.3–0.5 mm. |
Achenes | falling with scales, stramineous, rarely dark brown, ovoid to obpyriform, slightly to greatly compressed-trigonous, rarely thickly lenticular, angles prominent or abaxial angle obscure, 0.9–1.2 × 0.65–0.8 mm, apex narrowly to broadly truncate, neck short, often compressed more than body, at 20–30X finely rugulose with more than 20 horizontal ridges in a vertical series or reticulate or cancellate. |
with angles and longitudinal ridges ca. 6–10, rather prominent, broadly ovoid, less than 2 times longer then wide, (0.55–)0.65–0.75 × 0.3–0.4 mm, apex blunt, trabeculae distinct, 20–30. |
Tubercles | whitish to brown, pyramidal, lower than wide, often 3-lobed as viewed from the top, 0.1–0.3 × 0.4–0.65 mm. |
grayish, mostly appressed, pyramidal, often depressed, 0.1–0.2 × 0.1–0.25 mm. |
Eleocharis bolanderi |
Eleocharis bella |
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Phenology | Fruiting late spring–summer. | Fruiting spring–summer. |
Habitat | Fresh, often summer-dry meadows, springs, seeps, stream margins | Bare, often drying soil of stream alluvium, lake margins, wet meadows |
Elevation | 1000–3400 m (3300–11200 ft) | 200–2900 m (700–9500 ft) |
Distribution |
CA; CO; ID; NV; OR; UT
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AZ; CA; ID; MT; NM; NV; OR; WA; Mexico (Chihuahua)
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Discussion | Eleocharis bolanderi is clearly distinct from E. montevidensis, from which it differs in its dense, tufted habit with short, caudexlike rhizomes, its leaf sheaths without a tooth, its achene and tubercle shapes, and its acute floral scales. Specimens of Eleocharis bolanderi without rhizomes or achenes are easily confused with E. decumbens, which often may be distinguished by culms 0.5–2 mm wide, and spikelets with scales sometimes more than 3 mm long. The tubercles of E. bolanderi are usually poorly developed and much lower than wide; in E. decumbens they are usually well developed and about as high as wide. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Eleocharis bella and E. acicularis seem to be amply distinct; putative hybrids are unknown. The occasional plants of E. bella with evident rhizomes, which include the type, are otherwise identical to plants apparently without rhizomes. Eleocharis bella is very similar to E. cancellata. There is an Illinois collection from Peoria in 1901, from the alluvial banks of the Illinois River. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23. | FNA vol. 23, p. 110. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis | Cyperaceae > Eleocharis > subg. Scirpidium |
Sibling taxa | ||
Synonyms | E. montevidensis var. bolanderi | E. acicularis var. bella, E. acicularis var. minima |
Name authority | A. Gray: Proc. Amer. Acad. Arts 7: 392. (1868) | (Piper) Svenson: Rhodora 31: 201. (1929) |
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