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beautiful spikerush, delicate spikerush, pretty spikerush

California twisted spikerush, twist-stem spikerush

Habit Plants annual, rarely perennial, usually densely tufted; rhizomes rarely evident, 0.2–0.3 mm thick, internodes 1–5 mm, scales not evident. Plants perennial; rhizomes 1.5–2 mm thick, scales persistent, 7–9 mm, thinly papery, sometimes fibrous; resting buds unknown; caudices present, hard, 3 mm thick.
Culms

often ascending or spreading, 4-angled or sometimes terete, sometimes sulcate, 1–7 cm × 0.2–0.3 mm, soft to firm.

erect, markedly spirally twisted, markedly obliquely contracted near spikelet, when dry often with to 6 broad, rounded ridges on each side, greatly compressed, 3–4 times as wide as thick, 20–40 cm × 1.5–2.5 mm, firm;

culm tufts not proximally bulbous.

Leaves

sheaths stramineous, distal sheaths often splitting abaxially, slightly inflated distally, oblique, apex acute.

distal leaf sheaths stramineous to medium (or dark) brown, papery, apex often dark brown to reddish, broadly obtuse.

Spikelets

ovoid, 1.5–4 × 0.8–2 mm, apex acute;

floral scales 4–15, 8 per mm of rachilla, colorless or reddish brown, midrib region green, ovate-lanceolate, not folded lengthwise, 1–1.5 × 0.5–0.7 mm, mibrib obscure to somewhat keeled, apex narrowly acute to acuminate, slightly recurved.

6–8 × 2–3 mm;

proximal scale empty, 3–4 mm, shorter than to equaling spikelet;

floral scales 8–10 per spikelet, 3.5–5 × 2 mm.

Flowers

perianth bristles absent;

anthers 0.3–0.5 mm.

Perianth

bristles 0–5, unequal, rudimentary to equaling achene, the shorter stout, smooth or nearly so, the longer slender, densely spinulose.

Achenes

with angles and longitudinal ridges ca. 6–10, rather prominent, broadly ovoid, less than 2 times longer then wide, (0.55–)0.65–0.75 × 0.3–0.4 mm, apex blunt, trabeculae distinct, 20–30.

stramineous to medium brown, thickly trigonous, 1.75–2.75 × 1–1.25 mm;

beak 0.3–0.6 mm.

Tubercles

grayish, mostly appressed, pyramidal, often depressed, 0.1–0.2 × 0.1–0.25 mm.

0.25–0.6 × 0.3–0.55 mm.

Anthers

1.8 –3 mm.

Eleocharis bella

Eleocharis torticulmis

Phenology Fruiting spring–summer. Fruiting summer (Jun–Jul).
Habitat Bare, often drying soil of stream alluvium, lake margins, wet meadows Fens, wet meadows, vernal ponds
Elevation 200–2900 m (700–9500 ft) 1100 m (3600 ft)
Distribution
from FNA
AZ; CA; ID; MT; NM; NV; OR; WA; Mexico (Chihuahua)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Eleocharis bella and E. acicularis seem to be amply distinct; putative hybrids are unknown. The occasional plants of E. bella with evident rhizomes, which include the type, are otherwise identical to plants apparently without rhizomes. Eleocharis bella is very similar to E. cancellata. There is an Illinois collection from Peoria in 1901, from the alluvial banks of the Illinois River.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Eleocharis torticulmis is known only from the botanically rich Butterfly Valley in Plumas County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 110. FNA vol. 23, p. 115.
Parent taxa Cyperaceae > Eleocharis > subg. Scirpidium Cyperaceae > Eleocharis > subg. Zinserlingia
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms E. acicularis var. bella, E. acicularis var. minima
Name authority (Piper) Svenson: Rhodora 31: 201. (1929) S. G. Smith: Novon 11: 250, figs. 2, 4E–I. (2001)
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