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beautiful spikerush, delicate spikerush, pretty spikerush

four-angle spikerush, square-stem spike-rush, square-stem spikesedge

Habit Plants annual, rarely perennial, usually densely tufted; rhizomes rarely evident, 0.2–0.3 mm thick, internodes 1–5 mm, scales not evident. Plants perennial; rhizomes 1.5–4 mm thick, soft to hard, longer internodes 3–8 cm, scales 5–10 mm, tubers absent.
Culms

often ascending or spreading, 4-angled or sometimes terete, sometimes sulcate, 1–7 cm × 0.2–0.3 mm, soft to firm.

acutely quadrangular, (30–)45–105 cm × (1–)2–5.4 mm, soft to firm, internally spongy, transverse septa incomplete;

plants never forming filiform, flaccid culms.

Leaves

sheaths stramineous, distal sheaths often splitting abaxially, slightly inflated distally, oblique, apex acute.

distal leaf sheaths persistent, membranous, apex narrowly acute to acuminate, sometimes prolonged into a bladelike portion to 8 cm.

Spikelets

ovoid, 1.5–4 × 0.8–2 mm, apex acute;

floral scales 4–15, 8 per mm of rachilla, colorless or reddish brown, midrib region green, ovate-lanceolate, not folded lengthwise, 1–1.5 × 0.5–0.7 mm, mibrib obscure to somewhat keeled, apex narrowly acute to acuminate, slightly recurved.

not proliferous, (15–)20–76 × 3–5(–6) mm;

rachilla joints bearing obscure winglike remnants of floral scales;

proximal scale empty, amplexicaulous, (1–)2.2–5.4 mm;

floral scales (28–)45–135, 2–3 per mm of rachilla, stramineous to pale brown, usually with pale to dark brown submarginal band, midrib region sometimes greenish, broadly obovate to ovate, (4–)4.5–6.2 × 2.8–5 mm, subcartilaginous, apex rounded to obtuse.

Flowers

perianth bristles absent;

anthers 0.3–0.5 mm.

perianth bristles 6–7, whitish to brown, slender, often markedly unequal, shorter than achene or some equalling tubercle, sparsely retrorsely spinulose to smooth;

anthers stramineous to red-brown, 2.3–2.9 mm;

styles 3-fid, sometimes 2-fid.

Achenes

with angles and longitudinal ridges ca. 6–10, rather prominent, broadly ovoid, less than 2 times longer then wide, (0.55–)0.65–0.75 × 0.3–0.4 mm, apex blunt, trabeculae distinct, 20–30.

yellow or pale green to brown or purplish, biconvex, obovoid to obpyriform, 1.8–3 × 1.3–2 mm, almost smooth to markedly sculptured at 10–15X, each face with 19–38 rows of almost linear, transversely elongated cells, which are sometimes isodiametric at achene base, apex often constricted to neck 0.3–0.4 mm wide.

Tubercles

grayish, mostly appressed, pyramidal, often depressed, 0.1–0.2 × 0.1–0.25 mm.

dark brown or whitish, deltoid to high-pyramidal or lanceoloid, 0.7–1.5 × 0.4–1 mm, often spongy.

Eleocharis bella

Eleocharis quadrangulata

Phenology Fruiting spring–summer. Fruiting early summer–winter.
Habitat Bare, often drying soil of stream alluvium, lake margins, wet meadows Shallow water of fresh lake and pond shores, marshes
Elevation 200–2900 m (700–9500 ft) 10–600 m (0–2000 ft)
Distribution
from FNA
AZ; CA; ID; MT; NM; NV; OR; WA; Mexico (Chihuahua)
[WildflowerSearch map]
[BONAP county map]
from FNA
AR; CA; CT; DE; FL; GA; IL; IN; KY; LA; MA; MI; MO; MS; NC; NJ; NY; OH; OK; OR; PA; SC; TX; VA; WI; WV; ON; s to c Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Eleocharis bella and E. acicularis seem to be amply distinct; putative hybrids are unknown. The occasional plants of E. bella with evident rhizomes, which include the type, are otherwise identical to plants apparently without rhizomes. Eleocharis bella is very similar to E. cancellata. There is an Illinois collection from Peoria in 1901, from the alluvial banks of the Illinois River.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

We have not seen voucher specimens for the reports of Eleocharis quadrangulata from Kansas. Plants with greenish achenes, longer bristles, and longer anthers than the average are known from Tennessee.

The tubercles of Eleocharis quadrangulata are often spongy as in E. obtusetrigona.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 110. FNA vol. 23, p. 120.
Parent taxa Cyperaceae > Eleocharis > subg. Scirpidium Cyperaceae > Eleocharis > subg. Limnochloa
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms E. acicularis var. bella, E. acicularis var. minima Scirpus quadrangulatus, E. quadrangulata var. crassior, Scirpus albomarginatus, Scirpus marginatus
Name authority (Piper) Svenson: Rhodora 31: 201. (1929) (Michaux) Roemer & Schultes: in J. J. Roemer et al., Syst. Veg. 2: 155. (1817)
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