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beautiful spikerush, delicate spikerush, pretty spikerush

black sand spikerush, broad fruit spikerush

Habit Plants annual, rarely perennial, usually densely tufted; rhizomes rarely evident, 0.2–0.3 mm thick, internodes 1–5 mm, scales not evident. Plants perennial, forming dense clumps, often stoloniferous.
Rhizomes

present, caudexlike, mostly concealed by crowded culms, vertical or ascending, 2 mm thick;

internodes very short, scales disintegrating to fibers, 4 mm, papery.

Culms

often ascending or spreading, 4-angled or sometimes terete, sometimes sulcate, 1–7 cm × 0.2–0.3 mm, soft to firm.

erect to ascending or arching, acutely quadrangular, deeply sulcate, 7–50 cm × 0.3–0.5 mm, smooth, firm.

Leaves

sheaths stramineous, distal sheaths often splitting abaxially, slightly inflated distally, oblique, apex acute.

distal leaf sheaths persistent, proximally brown or reddish, distally stramineous to green, thickly membranous, apex acute or subacute.

Spikelets

ovoid, 1.5–4 × 0.8–2 mm, apex acute;

floral scales 4–15, 8 per mm of rachilla, colorless or reddish brown, midrib region green, ovate-lanceolate, not folded lengthwise, 1–1.5 × 0.5–0.7 mm, mibrib obscure to somewhat keeled, apex narrowly acute to acuminate, slightly recurved.

basal spikelets absent; often proliferous, ovoid, terete, 3–10 × 2–3 mm, apex acute;

proximal scale empty, amplexicaulous, similar to floral scales;

subproximal scale with a flower;

floral scales spiraled, 8–15, 4 per mm of rachilla, orange-brown, midrib region broadly stramineous or like flanks, ovate, 2–3 × 1.2–2 mm, membranous, midrib evident to prominent, apex acute (to rounded).

Flowers

perianth bristles absent;

anthers 0.3–0.5 mm.

perianth bristles 6 or fewer, stramineous, unequal, some or most equaling achene, spinules obscure, sparse, retrorse, sharply acute.

Stamens

3;

anthers 1.2–1.5 mm.

Styles

3-fid.

Achenes

with angles and longitudinal ridges ca. 6–10, rather prominent, broadly ovoid, less than 2 times longer then wide, (0.55–)0.65–0.75 × 0.3–0.4 mm, apex blunt, trabeculae distinct, 20–30.

stramineous, broadly obpyriform, equilaterally trigonous, angles evident, blunt, 0.9–1.1 × 0.7–0.9 mm, apex nearly truncate, smooth or obscurely papillose or rugulose at 20X.

Tubercles

grayish, mostly appressed, pyramidal, often depressed, 0.1–0.2 × 0.1–0.25 mm.

stramineous, pyramidal, trigonous, usually acute, proximally trilobed, the lobes decurrent on achene, 0.5–0.9 × 0.7–0.9 mm.

Eleocharis bella

Eleocharis pachycarpa

Phenology Fruiting spring–summer. Fruiting late spring–summer.
Habitat Bare, often drying soil of stream alluvium, lake margins, wet meadows Fresh shores, streambeds, groundwater seeps
Elevation 200–2900 m (700–9500 ft) 100–1400 m (300–4600 ft)
Distribution
from FNA
AZ; CA; ID; MT; NM; NV; OR; WA; Mexico (Chihuahua)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NV; South America (Argentina, Chile) [Introduced in North America; introduced in Australia]
[WildflowerSearch map]
[BONAP county map]
Discussion

Eleocharis bella and E. acicularis seem to be amply distinct; putative hybrids are unknown. The occasional plants of E. bella with evident rhizomes, which include the type, are otherwise identical to plants apparently without rhizomes. Eleocharis bella is very similar to E. cancellata. There is an Illinois collection from Peoria in 1901, from the alluvial banks of the Illinois River.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

H. K. Svenson (1957) placed Eleocharis pachycarpa in ser. Sulcatae Svenson. Contrary to his statement that perianth bristles are often lacking, they are present in all of the specimens we have seen. Superficially similar to E. bolanderi, from which it may readily be distinguished by its oblique leaf sheath summits, 4-angled culms, pyramidal tubercles, and often proliferating spikelets. The earliest North American specimen we have seen was collected in 1919 in Tuolumne County, California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 110. FNA vol. 23, p. 91.
Parent taxa Cyperaceae > Eleocharis > subg. Scirpidium Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms E. acicularis var. bella, E. acicularis var. minima
Name authority (Piper) Svenson: Rhodora 31: 201. (1929) E. Desvaux: in C. Gay, Fl. Chil. 6: 174. (1853)
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