Eleocharis bella |
Eleocharis occulta |
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beautiful spikerush, delicate spikerush, pretty spikerush |
limestone spike-rush |
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Habit | Plants annual, rarely perennial, usually densely tufted; rhizomes rarely evident, 0.2–0.3 mm thick, internodes 1–5 mm, scales not evident. | Plants perennial, densely cespitose; rhizomes concealed by persistent dead culm bases, often ascending, short, 3–5 mm thick, hard, cortex persistent, internodes very short, scales decaying to coarse fibers, 5–12 mm, papery. |
Culms | often ascending or spreading, 4-angled or sometimes terete, sometimes sulcate, 1–7 cm × 0.2–0.3 mm, soft to firm. |
subterete to slightly compressed, less than 2 times wider than thick, with 4–7 blunt ridges when dry, 27–56 cm × 0.2–0.5(–0.7) mm, firm to hard, spongy. |
Leaves | sheaths stramineous, distal sheaths often splitting abaxially, slightly inflated distally, oblique, apex acute. |
distal leaf sheaths persistent, not splitting, proximally red, distally green to stramineous, often inflated, thinly papery to membranous, sometimes translucent, apex broadly obtuse to subtruncate, often callose, tooth absent. |
Spikelets | ovoid, 1.5–4 × 0.8–2 mm, apex acute; floral scales 4–15, 8 per mm of rachilla, colorless or reddish brown, midrib region green, ovate-lanceolate, not folded lengthwise, 1–1.5 × 0.5–0.7 mm, mibrib obscure to somewhat keeled, apex narrowly acute to acuminate, slightly recurved. |
ovoid, 3–10 × 2–3 mm, apex acute; proximal scale amplexicaulous, apex 2-fid; subproximal scale with a flower; floral scales spreading in fruit, 20–50, 7 per mm of rachilla, medium brown, midrib region often narrowly stramineous, carinate, lanceolate-attenuate, 2–2.8 × 1 mm, apex 2-fid. |
Flowers | perianth bristles absent; anthers 0.3–0.5 mm. |
perianth bristles 3 or absent, stramineous to pale brown, rudimentary to 1/2 achene length, obscurely retrorsely spinulose; stamens 3; anthers orange-brown, 0.7–1.3 mm; styles 3-fid. |
Achenes | with angles and longitudinal ridges ca. 6–10, rather prominent, broadly ovoid, less than 2 times longer then wide, (0.55–)0.65–0.75 × 0.3–0.4 mm, apex blunt, trabeculae distinct, 20–30. |
falling with scales, medium or dark brown, obpyriform, nearly equilaterally obscurely trigonous or cross section nearly circular, 0.7–1 × 0.5–0.65 mm, neck distinct or rarely absent, obscurely rugulose at 10–30X, 30 or more low, blunt horizontal ridges in vertical series. |
Tubercles | grayish, mostly appressed, pyramidal, often depressed, 0.1–0.2 × 0.1–0.25 mm. |
brown, depressed-pyramidal, often rudimentary, 0.1–0.15 × 0.2 mm. |
Eleocharis bella |
Eleocharis occulta |
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Phenology | Fruiting spring–summer. | Fruiting spring (Mar–May), sometimes summer (Jul). |
Habitat | Bare, often drying soil of stream alluvium, lake margins, wet meadows | Seasonally wet, calcareous seeps, depressions, swales, rock crevises, rocky stream beds, stream banks, wet meadows, pond margins, often on limestone |
Elevation | 200–2900 m (700–9500 ft) | 80–300 m (300–1000 ft) |
Distribution |
AZ; CA; ID; MT; NM; NV; OR; WA; Mexico (Chihuahua)
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OK; TX |
Discussion | Eleocharis bella and E. acicularis seem to be amply distinct; putative hybrids are unknown. The occasional plants of E. bella with evident rhizomes, which include the type, are otherwise identical to plants apparently without rhizomes. Eleocharis bella is very similar to E. cancellata. There is an Illinois collection from Peoria in 1901, from the alluvial banks of the Illinois River. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Eleocharis occulta is very invariable in contrast to the extreme variability of E. compressa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 110. | FNA vol. 23, p. 83. |
Parent taxa | Cyperaceae > Eleocharis > subg. Scirpidium | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis |
Sibling taxa | ||
Synonyms | E. acicularis var. bella, E. acicularis var. minima | |
Name authority | (Piper) Svenson: Rhodora 31: 201. (1929) | S. G. Smith: Novon 11: 247, figs. 2, 3E–K. (2001) |
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