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purple spike-rush

California twisted spikerush, twist-stem spikerush

Habit Plants tufted, without creeping rhizomes. Plants perennial; rhizomes 1.5–2 mm thick, scales persistent, 7–9 mm, thinly papery, sometimes fibrous; resting buds unknown; caudices present, hard, 3 mm thick.
Culms

2–12(–19) cm × 0.2–0.4 mm.

erect, markedly spirally twisted, markedly obliquely contracted near spikelet, when dry often with to 6 broad, rounded ridges on each side, greatly compressed, 3–4 times as wide as thick, 20–40 cm × 1.5–2.5 mm, firm;

culm tufts not proximally bulbous.

Leaves

distal leaf sheaths firm, distally tightly sheathing, apex acute.

distal leaf sheaths stramineous to medium (or dark) brown, papery, apex often dark brown to reddish, broadly obtuse.

Spikelets

ovoid to ellipsoid, 2–6(–8) × 1–2.5 mm, apex acute;

proximal scale with or without flower, not amplexicaulous;

floral scales to 100, 15–19 per mm of rachilla, often loosely appressed, dark red-brown to stramineous, ovate to elliptic, 0.6–1.3 × 0.3–0.7 mm, membranous, apex rounded to acute.

6–8 × 2–3 mm;

proximal scale empty, 3–4 mm, shorter than to equaling spikelet;

floral scales 8–10 per spikelet, 3.5–5 × 2 mm.

Flowers

perianth bristles (0–)4–6, typically 4, colorless to whitish, vestigial to 1/2 as long as achene, smooth or spinuliferous;

styles 2-fid.

Perianth

bristles 0–5, unequal, rudimentary to equaling achene, the shorter stout, smooth or nearly so, the longer slender, densely spinulose.

Achenes

black, obovoid, biconvex, 0.3–0.5 × 0.3–0.4 mm, apex often constricted proximal to tubercle, smooth at 40X.

stramineous to medium brown, thickly trigonous, 1.75–2.75 × 1–1.25 mm;

beak 0.3–0.6 mm.

Tubercles

stramineous to whitish, umbonate to subconic, 0.1–0.2 × 0.1–0.2 mm, apex acute.

0.25–0.6 × 0.3–0.55 mm.

Anthers

1.8 –3 mm.

2n

= 20.

Eleocharis atropurpurea

Eleocharis torticulmis

Phenology Fruiting summer–fall (Jun–Sep). Fruiting summer (Jun–Jul).
Habitat Canal banks, hammocks, irrigation ditches, lake and pond margins, maritime shores, rice fields Fens, wet meadows, vernal ponds
Elevation 0–1800 m (0–5900 ft) 1100 m (3600 ft)
Distribution
from FNA
AL; CA; FL; GA; IA; KS; LA; MI; MO; NC; NE; NM; OK; SC; TX; WA; BC; Mexico; Central America; South America; West Indies; Europe (naturalized); Asia; Africa; Pacific Islands
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Eleocharis atropurpurea has been reported from Colorado, Montana, and Virginia; I have not seen voucher specimens.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Eleocharis torticulmis is known only from the botanically rich Butterfly Valley in Plumas County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 102. FNA vol. 23, p. 115.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Maculosae Cyperaceae > Eleocharis > subg. Zinserlingia
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms Scirpus atropurpureus
Name authority (Retzius) J. Presl & C. Presl: in C. B. Presl, Reliq. Haenk. 1: 196. (1828) S. G. Smith: Novon 11: 250, figs. 2, 4E–I. (2001)
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