Eleocharis atropurpurea |
Eleocharis pachycarpa |
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purple spike-rush |
black sand spikerush, broad fruit spikerush |
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Habit | Plants tufted, without creeping rhizomes. | Plants perennial, forming dense clumps, often stoloniferous. |
Rhizomes | present, caudexlike, mostly concealed by crowded culms, vertical or ascending, 2 mm thick; internodes very short, scales disintegrating to fibers, 4 mm, papery. |
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Culms | 2–12(–19) cm × 0.2–0.4 mm. |
erect to ascending or arching, acutely quadrangular, deeply sulcate, 7–50 cm × 0.3–0.5 mm, smooth, firm. |
Leaves | distal leaf sheaths firm, distally tightly sheathing, apex acute. |
distal leaf sheaths persistent, proximally brown or reddish, distally stramineous to green, thickly membranous, apex acute or subacute. |
Spikelets | ovoid to ellipsoid, 2–6(–8) × 1–2.5 mm, apex acute; proximal scale with or without flower, not amplexicaulous; floral scales to 100, 15–19 per mm of rachilla, often loosely appressed, dark red-brown to stramineous, ovate to elliptic, 0.6–1.3 × 0.3–0.7 mm, membranous, apex rounded to acute. |
basal spikelets absent; often proliferous, ovoid, terete, 3–10 × 2–3 mm, apex acute; proximal scale empty, amplexicaulous, similar to floral scales; subproximal scale with a flower; floral scales spiraled, 8–15, 4 per mm of rachilla, orange-brown, midrib region broadly stramineous or like flanks, ovate, 2–3 × 1.2–2 mm, membranous, midrib evident to prominent, apex acute (to rounded). |
Flowers | perianth bristles (0–)4–6, typically 4, colorless to whitish, vestigial to 1/2 as long as achene, smooth or spinuliferous; styles 2-fid. |
perianth bristles 6 or fewer, stramineous, unequal, some or most equaling achene, spinules obscure, sparse, retrorse, sharply acute. |
Stamens | 3; anthers 1.2–1.5 mm. |
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Styles | 3-fid. |
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Achenes | black, obovoid, biconvex, 0.3–0.5 × 0.3–0.4 mm, apex often constricted proximal to tubercle, smooth at 40X. |
stramineous, broadly obpyriform, equilaterally trigonous, angles evident, blunt, 0.9–1.1 × 0.7–0.9 mm, apex nearly truncate, smooth or obscurely papillose or rugulose at 20X. |
Tubercles | stramineous to whitish, umbonate to subconic, 0.1–0.2 × 0.1–0.2 mm, apex acute. |
stramineous, pyramidal, trigonous, usually acute, proximally trilobed, the lobes decurrent on achene, 0.5–0.9 × 0.7–0.9 mm. |
2n | = 20. |
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Eleocharis atropurpurea |
Eleocharis pachycarpa |
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Phenology | Fruiting summer–fall (Jun–Sep). | Fruiting late spring–summer. |
Habitat | Canal banks, hammocks, irrigation ditches, lake and pond margins, maritime shores, rice fields | Fresh shores, streambeds, groundwater seeps |
Elevation | 0–1800 m (0–5900 ft) | 100–1400 m (300–4600 ft) |
Distribution |
AL; CA; FL; GA; IA; KS; LA; MI; MO; NC; NE; NM; OK; SC; TX; WA; BC; Mexico; Central America; South America; West Indies; Europe (naturalized); Asia; Africa; Pacific Islands
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CA; NV; South America (Argentina, Chile) [Introduced in North America; introduced in Australia]
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Discussion | Eleocharis atropurpurea has been reported from Colorado, Montana, and Virginia; I have not seen voucher specimens. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
H. K. Svenson (1957) placed Eleocharis pachycarpa in ser. Sulcatae Svenson. Contrary to his statement that perianth bristles are often lacking, they are present in all of the specimens we have seen. Superficially similar to E. bolanderi, from which it may readily be distinguished by its oblique leaf sheath summits, 4-angled culms, pyramidal tubercles, and often proliferating spikelets. The earliest North American specimen we have seen was collected in 1919 in Tuolumne County, California. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 23, p. 102. | FNA vol. 23, p. 91. |
Parent taxa | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Maculosae | Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Tenuissimae |
Sibling taxa | ||
Synonyms | Scirpus atropurpureus | |
Name authority | (Retzius) J. Presl & C. Presl: in C. B. Presl, Reliq. Haenk. 1: 196. (1828) | E. Desvaux: in C. Gay, Fl. Chil. 6: 174. (1853) |
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