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purple spike-rush

decumbent spikerush

Habit Plants tufted, without creeping rhizomes. Plants perennial, densely tufted; rhizomes often hidden by culms and roots, fairly long, 3–4 mm thick, hard, cortex persistent, longer internodes from very short to 5 mm, scales usually clearly evident, disintegrating to fibers, 20–25 mm, papery.
Culms

2–12(–19) cm × 0.2–0.4 mm.

terete, often with 10–18 blunt ridges when dry, 10–50 cm × 0.3–2 mm, firm to rigid, spongy.

Leaves

distal leaf sheaths firm, distally tightly sheathing, apex acute.

distal leaf sheaths persistent, not splitting, proximally brown or reddish, distally stramineous, brown, reddish or green, often inflated, papery, apex mostly dark red-brown, subtruncate to obtuse, often callose, tooth absent.

Spikelets

ovoid to ellipsoid, 2–6(–8) × 1–2.5 mm, apex acute;

proximal scale with or without flower, not amplexicaulous;

floral scales to 100, 15–19 per mm of rachilla, often loosely appressed, dark red-brown to stramineous, ovate to elliptic, 0.6–1.3 × 0.3–0.7 mm, membranous, apex rounded to acute.

ovoid, 3–8 × 2–2.5 mm, apex acute;

proximal scale amplexicaulous, entire;

subproximal scale empty or with flower;

floral scales appressed in fruit, 10–20, 3 per mm of rachilla, orange-brown, midrib regions often greenish, ovate, 3–3.5 × 1.5 mm, apex entire, acute, often carinate in distal part of spikelet.

Flowers

perianth bristles (0–)4–6, typically 4, colorless to whitish, vestigial to 1/2 as long as achene, smooth or spinuliferous;

styles 2-fid.

perianth bristles 6, stramineous, stout, nearly equal, mostly equaling or exceeding tubercle, (0.5–)1–2.2 mm, prominently retrorsely spinulose;

stamens 3(?);

anthers dark yellow to stramineous, 1.2–1.5;

styles 3-fid.

Achenes

black, obovoid, biconvex, 0.3–0.5 × 0.3–0.4 mm, apex often constricted proximal to tubercle, smooth at 40X.

falling with scales, dark brown, obpyriform, nearly eqilaterally- to greatly compressed-trigonous, angles slightly prominent, 1–1.3 × 0.75–0.9 mm, neck absent or short, finely rugulose at 20X with more than 20 horizontal ridges in vertical series, or reticulate or cancellate at 20–30X.

Tubercles

stramineous to whitish, umbonate to subconic, 0.1–0.2 × 0.1–0.2 mm, apex acute.

well developed, whitish, pyramidal, as high as wide to much lower than wide, 0.2–0.6 × 0.4–0.7 mm.

2n

= 20.

Eleocharis atropurpurea

Eleocharis decumbens

Phenology Fruiting summer–fall (Jun–Sep). Fruiting summer–fall.
Habitat Canal banks, hammocks, irrigation ditches, lake and pond margins, maritime shores, rice fields Wet fresh meadows, seeps, and lakeshores, in interior montane conifer forests and alpine zones
Elevation 0–1800 m (0–5900 ft) 700–3500 m (2300–11500 ft)
Distribution
from FNA
AL; CA; FL; GA; IA; KS; LA; MI; MO; NC; NE; NM; OK; SC; TX; WA; BC; Mexico; Central America; South America; West Indies; Europe (naturalized); Asia; Africa; Pacific Islands
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[BONAP county map]
from FNA
CA; OR
[BONAP county map]
Discussion

Eleocharis atropurpurea has been reported from Colorado, Montana, and Virginia; I have not seen voucher specimens.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eleocharis decumbens is known only from Shasta and Tulare counties in California.

Although Eleocharis decumbens has long been ignored or treated as a variety of E. montevidensis, it clearly is a very distinct species, from which it differs especially in its thick rhizomes with fibrous scales and its acute floral scales.

Specimens of Eleocharis decumbens without rhizomes or achenes are easily confused with the apparently closely related E. bolanderi, which often may be distinguished by culms no more than 0.5 mm wide and spikelets with scales no more than 3 mm long. The tubercles of E. decumbens are usually well differentiated from the achenes and about as high as wide; the tubercles of E. bolanderi are often poorly developed and much lower than wide. Three collections from Jackson and Klamath counties in Oregon, lack achenes but are probably E. decumbens.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 102. FNA vol. 23.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Maculosae Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms Scirpus atropurpureus E. montevidensis var. decumbens
Name authority (Retzius) J. Presl & C. Presl: in C. B. Presl, Reliq. Haenk. 1: 196. (1828) C. B. Clarke: Bull. Misc. Inform., addit. ser. 8: 23. (1908)
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