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purple spike-rush

glades spikerush

Habit Plants tufted, without creeping rhizomes. Plants perennial, densely cespitose; rhizomes concealed by persistent culm bases, short, 4–5 mm thick, hard, cortex persistent, internodes crowded, scales decaying to coarse fibers, 1 cm, papery.
Culms

2–12(–19) cm × 0.2–0.4 mm.

greatly compressed, 4–10 times wider than thick, often with 1 or 2 sharp ridges on 1 side, (8–)20–35 cm × 0.7–2.3 mm, hard, finely striate, spongy.

Leaves

distal leaf sheaths firm, distally tightly sheathing, apex acute.

distal leaf sheaths persistent, not splitting, proximally red or stramineous, distally green to stramineous, inflated, papery, apex dark brown, broadly obtuse to subtruncate, callose, tooth absent.

Spikelets

ovoid to ellipsoid, 2–6(–8) × 1–2.5 mm, apex acute;

proximal scale with or without flower, not amplexicaulous;

floral scales to 100, 15–19 per mm of rachilla, often loosely appressed, dark red-brown to stramineous, ovate to elliptic, 0.6–1.3 × 0.3–0.7 mm, membranous, apex rounded to acute.

ovoid, 4–9 × 2.5–4 mm, apex acute;

proximal scale clasping 2/3–3/4 of culm, apex 2-fid;

subproximal scale empty or with a flower;

floral scales spreading in fruit, 30–60, 6–9 per mm of rachilla, medium or pale brown, midrib region often paler, ovate-lanceolate, 2.5–3.5 × 1.5 mm, apex 2-fid, carinate in distal part of spikelet.

Flowers

perianth bristles (0–)4–6, typically 4, colorless to whitish, vestigial to 1/2 as long as achene, smooth or spinuliferous;

styles 2-fid.

perianth bristles 0(–5), stramineous to pale brown, 1/2 of to equaling achene length;

stamens 3;

anthers orange-brown, 0.7–1.5 mm;

styles 3-fid.

Achenes

black, obovoid, biconvex, 0.3–0.5 × 0.3–0.4 mm, apex often constricted proximal to tubercle, smooth at 40X.

falling with scales, yellow-brown to medium brown, obovoid to obpyriform, nearly equilaterally trigonous, cross section circular to slightly compressed, angles obscure or evident, 0.9–1.1 × 0.6–0.75, neck usually very short, finely rugulose at 10–30X, 20 or more low, blunt horizontal ridges in vertical series.

Tubercles

stramineous to whitish, umbonate to subconic, 0.1–0.2 × 0.1–0.2 mm, apex acute.

brown, depressed-pyramidal, often rudimentary, 0.1–0.25 × 0.2–0.3 mm.

2n

= 20.

Eleocharis atropurpurea

Eleocharis bifida

Phenology Fruiting summer–fall (Jun–Sep). Fruiting spring–summer (May–Jun).
Habitat Canal banks, hammocks, irrigation ditches, lake and pond margins, maritime shores, rice fields Seasonally wet places on limestone, cedar (juniper) glades, stream beds, prairies, ditches
Elevation 0–1800 m (0–5900 ft) 200–300 m (700–1000 ft)
Distribution
from FNA
AL; CA; FL; GA; IA; KS; LA; MI; MO; NC; NE; NM; OK; SC; TX; WA; BC; Mexico; Central America; South America; West Indies; Europe (naturalized); Asia; Africa; Pacific Islands
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[BONAP county map]
from FNA
AL; GA; KY; TN
[BONAP county map]
Discussion

Eleocharis atropurpurea has been reported from Colorado, Montana, and Virginia; I have not seen voucher specimens.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eleocharis bifida was previously included in E. compressa. A very few specimens of E. compressa var. compressa from the states where E. bifida is known are like E. bifida except for their entire proximal floral scales and evident rhizomes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 102. FNA vol. 23, p. 85.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleogenus > ser. Maculosae Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis
Sibling taxa
E. acicularis, E. aestuum, E. albida, E. ambigens, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Synonyms Scirpus atropurpureus
Name authority (Retzius) J. Presl & C. Presl: in C. B. Presl, Reliq. Haenk. 1: 196. (1828) S. G. Smith: Novon 11: 243, figs. 1F–J, 2. (2001)
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