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white spikerush

ambiguous spikesedge, éléocharide ambiquë

Habit Plants perennial; rhizomes soft, longer internodes 2–4 cm, cortex loose, scales fugaceous, 6 mm, thinly membranous and translucent. Plants perennial, mat-forming; rhizomes evident, long, 1–2 mm thick, firm, cortex persistent, longer internodes 1–3 cm, scales persistent, 5–10 mm, membranous, not fibrous.
Culms

not rooting at tips, terete, 10–40 cm, soft to firm, smooth.

terete, often with to 16 blunt ridges when dry, 25–80 cm × 0.5–1.5 mm, firm, internally spongy.

Leaves

distal leaf sheaths proximally brownish or sometimes reddish, distally stramineous to green.

distal leaf sheaths persistent, not splitting, proximally dark red, distally red to stramineous or green, often callose, thinly papery (to membranous), apex dark red-brown, obtuse to subtruncate, tooth sometimes present on some culms, to 0.3 mm.

Spikelets

ovoid to oblong-subcylindric, 4–12 × 2–3.5 mm, apex acute to rounded;

proximal scale empty, clasping 1/2 of culm, like floral scales;

subproximal 1 or 2 scales often empty;

floral scales 20–100, 10 per mm of rachilla, entirely stramineous or sometimes red-brown, ovate, (1.5–)2–2.5 × 1.5 mm, apex broadly rounded, entire.

ovoid to lanceoloid, 5–23 × 2–3(–4) mm, apex acute (to obtuse);

proximal scale amplexicaulous, entire;

subproximal scale with flower;

floral scales often spreading in fruit, 10–60, 3–4 per mm of rachilla, medium brown, rarely red-brown, midrib regions mostly stramineous to green, in proximal part of spikelet ovate, in distal part lanceolate, 2.5–3.5 × 1.7 mm, entire, apex acute to rarely obtuse, often carinate in distal part of spikelet.

Flowers

perianth bristles (5–)6(–8), brown, stout, the longer equaling achene or tubercle, retrorsely spinulose;

stamens 3;

anthers brown, 1 mm.

perianth bristles 2–4(–5), brown, slender to stout, usually very unequal, rudimentary to equaling achene;

stamens 3;

anthers brown, 1.5–2 mm;

styles 2-fid, sometimes some 3-fid.

Achenes

falling with scales, obovoid, angles keeled, 0.8–1 × 0.7–0.8 mm, apex with short neck.

not persistent, dark yellow or stramineous, obovoid to obpyriform, all biconvex or some (very rarely all) compressedtrigonous, angles obscure, 1–1.5 × 0.9–1.25 mm, apex rounded, neck absent (to very short), finely rugulose at 10–20X, with 20 or more horizontal ridges in vertical series and/or finely cancellate at 10–30X.

Tubercles

whitish to brown, mammillate to pyramidal, 0.2–0.3 × 0.3–0.35 mm, 1/3 or less as wide as achene.

brown or proximally whitish, pyramidal, depressed, rarely some as high as wide, 0.15–0.3(–0.5) × (0.3–)0.4–0.6 mm.

2n

= 44, 45, 46.

Eleocharis albida

Eleocharis ambigens

Phenology Fruiting summer. Fruiting spring–summer.
Habitat Coastal saltmarsh edges, sloughs, beaches, dune depressions, ditches Coastal (rarely inland) fresh to brackish pond shores, marshes, disturbed places
Elevation 0 m (0 ft) 0–10 m (0–0 ft)
Distribution
from FNA
AL; FL; GA; LA; MD; MS; NC; NM; SC; TX; Mexico; Bermuda
[WildflowerSearch map]
[BONAP county map]
from FNA
CT; FL; GA; LA; MA; MS; NC; NJ; RI; SC; TX; VA; QC
Discussion

In most spikelets, the bright brown stigmas contrast strikingly with the stramineous floral scales. We have not seen vouchers for H. K. Svenson’s (1937) reports of Eleocharis albida from Virginia. The collections we have seen from Maryland are from the 1800s.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Eleocharis ambigens is usually treated as a synonym of E. fallax because some specimens from Massachusetts to Georgia have some 3-fid styles and some obscurely trigonous achenes. A specimen of E. ambigens from Louisiana has trigonous achenes. Eleocharis ambigens is very similar to and perhaps better included in E. uniglumis (S.-O. Strandhede 1967). Intermediates with E. macrostachya occur in Louisiana and Texas. The depressed tubercles are diagnostic of E. ambigens. Eleocharis ambigens is sometimes confused with E. montevidensis, which is readily distinguished by its floral scales with broadly rounded apices.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 23, p. 99. FNA vol. 23, p. 78.
Parent taxa Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Albidae Cyperaceae > Eleocharis > subg. Eleocharis > sect. Eleocharis > ser. Eleocharis
Sibling taxa
E. acicularis, E. aestuum, E. ambigens, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
E. acicularis, E. aestuum, E. albida, E. atropurpurea, E. baldwinii, E. bella, E. bernardina, E. bicolor, E. bifida, E. bolanderi, E. brachycarpa, E. brittonii, E. cancellata, E. cellulosa, E. coloradoensis, E. compressa, E. cylindrica, E. decumbens, E. diandra, E. elliptica, E. elongata, E. engelmannii, E. equisetoides, E. erythropoda, E. fallax, E. flavescens, E. geniculata, E. intermedia, E. interstincta, E. kamtschatica, E. lanceolata, E. macrostachya, E. mamillata, E. melanocarpa, E. microcarpa, E. minima, E. montana, E. montevidensis, E. nana, E. nigrescens, E. nitida, E. obtusa, E. obtusetrigona, E. occulta, E. ovata, E. pachycarpa, E. palustris, E. parishii, E. parvula, E. quadrangulata, E. quinqueflora, E. radicans, E. ravenelii, E. retroflexa, E. reverchonii, E. robbinsii, E. rostellata, E. suksdorfiana, E. tenuis, E. torticulmis, E. tortilis, E. tricostata, E. tuberculosa, E. uniglumis, E. vivipara, E. wolfii
Name authority Torrey: Ann. Lyceum Nat. Hist. New York. 3: 304. (1836) Fernald: Rhodora 37: 394. (1935)
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