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California waterwort

mud-purslane, waterwort, élatine

Habit Herbs, submersed or growing on exposed but wet substrates, 1–5 cm. Herbs, submersed or emergent aquatic, glabrous.
Stems

decumbent to erect, branched.

erect, ascending, decumbent, or prostrate, with longitudinal air spaces, rooting at nodes.

Leaves

light green to green, sometimes becoming reddish in terrestrial plants;

stipules lanceolate, 0.5–0.6 mm, margins dentate, apex acute;

petiole 1–4 mm;

blade obovate to oblanceolate, 4–12(–15) × 1.2–3 mm, base narrowly cuneate, apex acute to obtuse.

stipules membranous;

petiole present or absent;

blade margins entire, with hydathodes.

Inflorescences

flowers usually solitary, sometimes 2[–3] per node.

Pedicels

1.5–2.5(–3.5) mm, recurved in fruit.

present or absent.

Flowers

sepals 4, usually equal, sometimes 1 reduced, oblong-ovate or widely lanceolate, 0.5–0.6 × 0.3–0.4 mm;

petals 4, greenish white, slightly reddish, or pink, elliptic or ovate, 1–1.5 × 0.5–0.6 mm;

stamens 8;

styles 4.

sepals 2–4, connate basally, equal or 1 smaller, not carinate, apex obtuse;

petals (0 or) 2–4, apex obtuse;

stamens [0–]1–8;

pistil 2–4-carpellate, ovary 2–4-locular, apex truncate;

styles 2–4;

stigmas 2–4.

Capsules

depressed-ovoid, 4-locular, 1.3–2.5 mm diam.

membranous.

Seeds

2–5 per locule, oblong to ellipsoid, curved 90–180°, 0.6–1 × 0.2 mm;

pits elliptic, length 1–3 times width, in 6–10 rows, (16–)20–29 per row.

2–33[–44] per locule, brown to yellowish brown, straight or curved (nearly circular in E. californica), surface with hexagonal, rectangular, elliptic, or ± round pits (pits oriented with longer dimension at right angles to length of seed).

x

= 9.

Elatine californica

Elatine

Phenology Flowering summer.
Habitat Pools, pond shores, rice fields, stream banks.
Elevation 0–1900(–2600) m. (0–6200(–8500) ft.)
Distribution
from FNA
AZ; CA; ID; MT; NM; NV; OR; UT; WA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from USDA
North America; Mexico; South America; Europe; Asia; Africa; Pacific Islands; Australia
[BONAP county map]
Discussion

Elatine californica is distinguished from other species of the genus by having long pedicels that are recurved in fruit and strongly curved seeds that can be nearly circular.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species ca. 25 (10 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals 2; capsules 2-locular.
E. minima
1. Petals usually 3–4, sometimes 0; capsules 3–4-locular.
→ 2
2. Sepals 4; petals 4; capsules 4-locular.
→ 3
3. Seeds curved 90–180°; pedicels 1.5–2.5(–3.5) mm, recurved in fruit; w United States.
E. californica
3. Seeds straight or curved to 15º; pedicels 0.1–23 mm, erect; Quebec.
E. ojibwayensis
2. Sepals 2–3 (rarely 4 in E. triandra); petals 3 (sometimes 0 in E. ambigua); capsules 3-locular.
→ 4
4. Pedicels recurved in fruit, 0.5–2.5 mm.
E. ambigua
4. Pedicels erect, 0–0.5 mm.
→ 5
5. Stamens 1–6, number variable within plant.
E. heterandra
5. Stamens 3.
→ 6
6. Seed pits ± round, (9–)14–17 per row.
E. brachysperma
6. Seed pits angular-hexagonal, (13–)16–35 per row.
→ 7
7. Seeds ellipsoid, pit length 3–5 times width; stipule margins entire.
E. chilensis
7. Seeds oblong or slenderly cylindric, pit length 1–3 times width; stipule margins dentate.
→ 8
8. Leaves reddish green; seed pit length 1–2 times width.
E. rubella
8. Leaves light green to green; seed pit length 2–3 times width.
→ 9
9. Leaf blades linear, lanceolate or narrowly oblong, apices acute or obtuse.
E. triandra
9. Leaf blades obovate or broadly spatulate, apices rounded to shallowly emarginate.
E. americana
Source FNA vol. 12, p. 350. FNA vol. 12, p. 349. Authors: Hamid Razifard, Gordon C. Tucker, Donald H. Les.
Parent taxa Elatinaceae > Elatine Elatinaceae
Sibling taxa
E. ambigua, E. americana, E. brachysperma, E. chilensis, E. heterandra, E. minima, E. ojibwayensis, E. rubella, E. triandra
Subordinate taxa
E. ambigua, E. americana, E. brachysperma, E. californica, E. chilensis, E. heterandra, E. minima, E. ojibwayensis, E. rubella, E. triandra
Synonyms E. californica var. williamsii, E. williamsii Crypta
Name authority A. Gray: Proc. Amer. Acad. Arts 13: 361, 364. (1878) Linnaeus: Sp. Pl. 1: 367. (1753): Gen. Pl. ed. 5, 172. (1754)
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