Echinocereus triglochidiatus
Echinocereus papillosus
claret-cup cactus, claretcup, kingcup cactus, Mojave mound cactus
allicoche cactus, allicoche hedgehog cactus, yellow-flower alicoche
usually erect or nearly so, cylindric (spheric), (2–)5–70 × (3–)5–13 cm;
ribs 5–8 or 8–12, crests slightly undulate (localized populations contain plants with strongly interrupted ribs);
areoles 10–40 mm apart.
mostly nearly erect when young, frequently decumbent in old age, cylindric, 4–15 × 2–3(–7) cm;
ribs (6–)7–10, crests strongly undulate;
areoles 10–15 mm apart.
(0–)3–11 per areole, straight to curved or contorted, appressed (radial spines) or spreading to projecting outward (some radials and central spines when present), white to yellow, gray, or black;
radial spines (0–)1–10 per areole, (0–)15–90 mm;
central spines 0–1(–4) per areole, angular, (0–)50–120 mm.
8–12 per areole, straight, appressed-spreading (radial spines) or projecting to appressed (central spines), white, yellow, or brown and white;
radial spines 7–10 per areole, to 15 mm;
central spines 1(–4) per areole, brown, terete, to 25 mm.
(4–)5–10 × 3–7 cm;
flower tube 20–35 mm;
flower tube hairs 1–2 mm;
inner tepals bright orange-red to dark red, proximally paler (bases sometimes yellow or white), (18–)25–40 × (5–)10–15 mm, tips thick and rigid;
anthers usually pink to purple;
nectar chamber 5–11 mm.
funnelform, 7–9 × 7–12 cm;
flower tube 20–30 × 10–30 mm;
flower tube hairs 1 mm;
inner tepals yellow, proximally orange-red to purple, 15–40 × 10–20 mm, tips relatively thin and delicate;
filaments yellowish to orange-red;
anthers yellow;
nectar chamber 4–6 mm.
green to yellow-green or pink (rarely red), (15–)20–35 mm, pulp white.
green, 15–25 mm, pulp white.
= 22.
= 22.
Echinocereus triglochidiatus
Echinocereus papillosus
Echinocereus triglochidiatus is the earliest name for a large group of diploid and polyploid taxa treated as conspecific by L. D. Benson (1969, 1982). The tetraploids are now recognized separately as E. coccineus, including E. polyacanthus Engelmann of Mexico. The diploids in the flora area are here divided into two allopatric species: E. triglochidiatus and E. arizonicus.
Plants in the western portion of the range of Echinocereus triglochidiatus have been called E. triglochidiatus var. mojavensis (Engelmann & J. M. Bigelow) L. D. Benson. That taxon includes curly-spined plants (mainly in California) and straight-spined plants (including most populations in Arizona, Utah, and western Colorado). The latter were mapped by L. D. Benson (1969, 1982) as part of his concept of E. triglochidiatus var. melanacanthus. Plants with the fewest and largest spines, called E. triglochidiatus var. triglochidiatus, occupy the eastern portion of the species’ distribution. The largest spines, whether central or radial, of var. triglochidiatus are sharply angular in cross section and 1–2 mm thick.
Less distinctive plants have been called Echinocereus triglochidiatus var. gonacanthus, a name carelessly applied to miscellaneous plants throughout the range of var. triglochidiatus. A well-known population at White Sands, New Mexico, inhabits saline flats adjacent to pure gypsum dunes. The unusually large, southernmost plants at White Sands, New Mexico, shrink to the same size as northern plants when grown together in a common garden (D. Weniger 1970). The epithet inermis has been applied at various taxonomic ranks to individual plants with spines absent or nearly so in the eastern portion of var. mojavensis (in and around southeastern Utah).
A geographically distant tetraploid, Echinocereus coccineus var. paucispinus, superficially resembles some eastern E. triglochidiatus but may be distinguished by its relatively terete and more consistently straight spines. Small plants of var. triglochidiatus without reproductive structures might be confused with E. fendleri; records of E. triglochidatus from near the Mexican border probably are misidentifications of E. fendleri in vegetative condition.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Small-stemmed plants were designated as Echinocereus papillosus var. angusticeps (Clover) W. T. Marshall, reportedly endemic near Linn, Texas. The status of the proposed variety remains unresolved but appears weak; W. Blum et al. (1998) and E. F. Anderson (2001) considered the name to be a synonym of typical E. papillosus.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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