Echinocereus enneacanthus |
Echinocereus bonkerae |
|||||
---|---|---|---|---|---|---|
alicoche, Mexican strawberry pitaya, pitaya, smallspine pitaya, strawberry cactus |
bonker hedgehog, pinkflower hedgehog cactus |
|||||
Habit | Plants branched forming dense or lax clumps with 20–100(–500) branches, usually branching before flowering. | Plants 5–35-branched, ultimately forming clumps, branching at or before sexual maturation. | ||||
Stems | some-what lax often sprawling, longest stems sometimes prostrate, cylindric, 8–40(–100?) × 3.2–15 cm; ribs (6–)7–10(–12), crests essentially uninterrupted; areoles (11–)14–52 mm apart. |
mostly erect, ovoid to cylindric, 15–30(–50) × 3.5–7 cm; ribs (11–)12–18(–20), crests slightly undulate; areoles 8–20 mm apart. |
||||
Spines | 6–14 per areole, straight or central spines slightly curved throughout their lengths, ± opaque, white, pale tan, or purplish gray, often extensively tipped or banded with brown; radial spines 5–10(–13) per areole, 9.5–40(–47) mm, usually less than 1/2 as long as central spines; central spines 1–4(–5) per areole, all or mostly projecting, abaxial spine porrect or descending, frequently compressed or angular in cross section (sometimes sulcate, keeled, or striate), (12–)20–84(–96) mm. |
(9–)12–17 per areole, straight in short-spined forms, sometimes curved or twisted in long-spined forms, appressed (radial spines) or porrect to spreading or descending (central spines when present), dull yellowish to brown or white to gray, often brown to black especially at bases or tips; radial spines (9–)11–16 per areole, 5–18 mm; central spines (0–)1(–3) per areole, 2–100 mm, white to yellow or brown, often becoming gray, all terete. |
||||
Flowers | (4.5–)5–7.5 × 5–5.6(–9) cm; flower tube 10–30 × 10–22(–40) mm; flower tube hairs 1–2 mm; inner tepals pink or magenta, darkest proximally, 28–55 × 8–14(–20) mm, tips relatively thin and delicate; anthers yellow; nectar chamber 4–6 mm. |
5–7 × 4–9 cm; flower tube 12–20 × 10–30 mm; flower tube hairs 1 mm; inner tepals deep magenta (to dark purple) with darker midstripes, proximally green (to very dark purple), 20–60 × 8.5–20 mm, tips relatively thin and delicate; anthers yellow; nectar chamber 2–4 mm. |
||||
Fruits | pale yellow-green or dull reddish, 20–30 mm, pulp white or pale pink. |
bright red or orange red, 15–25 mm, pulp white (or pale pink). |
||||
2n | = 22. |
= 22. |
||||
Echinocereus enneacanthus |
Echinocereus bonkerae |
|||||
Phenology | Flowering Mar–May; fruiting 2 months after flowering. | |||||
Habitat | Sonoran Desert upland, interior chaparral, desert grasslands, pinyon-juniper woodlands | |||||
Elevation | 700-2000 m (2300-6600 ft) | |||||
Distribution |
TX; Mexico
|
AZ
|
||||
Discussion | Varieties 2 (2 in the flora). The commonly recognized concept of Echinocereus enneacanthus var. enneacanthus (W. O. Moore 1967; D. Weniger 1970; L. D. Benson 1982) pertained to the small eastern var. brevispinus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Plants in most populations of Echinocereus bonkerae have radial spines almost pectinately arranged and central spines very short or absent, creating the superficial appearance of E. dasyacanthus, E. pectinatus, or similar species. Formerly, all short-spined plants in central and southern Arizona were treated as E. bonkerae, including unusually short-spined individuals of E. fendleri and E. fasciculatus. Today we recognize that E. bonkerae, E. fendleri, and E. fasciculatus all vary from short-spined to long-spined. Populations at the lowest altitude for the species have taller stems and unusually long, slender central spines (to 10 cm); they have recently been named Echinocereus apachensis. Such plants were part of the basis for L. D. Benson’s polyphyletic concept (1969, 1982) of E. fasciculatus var. boyce-thompsonii (see 8. E. fasciculatus). Echinocereus bonkerae is a poorly defined species, frequently lumped with E. fendleri or E. fasciculatus, and it is not always identifiable in the field. Although rib number, blooming season, habitat preference, and, in particular regions, spine length or central spine presence are helpful characteristics, ploidy level is the most objective criterion for distinguishing E. bonkerae from E. fasciculatus. Immature plants of E. bonkerae are like E. fasciculatus and unlike the immature plants of E. fendleri, which are tuberculate with their spines long, few, soft, and often curved or twisted. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
||||
Key |
|
|||||
Source | FNA vol. 4, p. 162. | FNA vol. 4. | ||||
Parent taxa | Cactaceae > subfam. Cactoideae > Echinocereus | Cactaceae > subfam. Cactoideae > Echinocereus | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | E. apachensis, E. fasciculatus var. bonkerae, E. fendleri var. bonkerae | |||||
Name authority | Engelmann: in F. A. Wislizenus, Mem. Tour N. Mexico, 111. (1848) | Thornber & Bonker: Fantast. Clan, 71, 85, plates opposite 23, 72. (1932) | ||||
Web links |