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biznaga-tonel manca caballo, devil's pincushion, devil's-head, horse-crippler, manca caballo

biznaga-tonelmancamula, blue barrel cactus, devilshead, silverbell cactus, Turk's head cactus, visnaga meloncillo

Habit Plants unbranched (very rarely branched). Plants normally unbranched.
Stems

pale gray-green (desert populations) to grass green (eastern populations), above-ground portion flat-topped, hemispheric in old age but usually deep-seated, flush with soil surface, 10–30 × 10–30 cm;

ribs 13–27, very prominent, straight, vertical, or sinuous on desiccated plants, crests ± sharp, without depressions between areoles but sometimes areoles recessed part way into rib.

pale gray-green to bright gray-blue, flat-topped or hemispheric and deep-seated in sub-strate, spheric with age or stoutly short cylindric (remaining hemispheric at high elevations), 4–25(–45) × 8–15(–20) cm;

ribs (7–)8(–9), vertical to helically curving around stem, rib crests broadly rounded, uninterrupted or slightly constricted between areoles.

Spines

(6–)7–8 per areole, mostly decurved or 1 porrect and straight, pale tan, pink, reddish to gray, terete to flattened, annulate, not hiding stem surfaces, minutely canescent with laterally compressed unicellular trichomes;

radial spines (5–)6–7 per areole;

central spine 1 per areole, porrect or descending, straight or distally decurved, (20–)40–60(–80) × 1.5–4(–8) mm.

(5–)8(–10) per areole, loosely projecting or strongly decurved, pink, gray, tan, or brown, strongly annulate-ridged, subulate, ± flattened, glabrous, generally not hiding stem surface;

radial spines 5(–8) per areole, similar to central spines;

central spines 1(–3) per areole, 18–43 × 1–2.5(–3) mm, longest spine usually descending, straight or decurved throughout its length.

Flowers

5–6 × 5–6 cm;

inner tepals bright rose-pink to pale silvery-pink, proximally orange to red, (15–)28–32 × (3–)6(–9) mm, margins usually erose;

stigma lobes pink to pinkish white.

5–7 × 5–6.5(–9.5) cm;

inner tepals bright rose-pink or magenta, color ± uniform from base to apex, 3 × 1.5 cm, margins entire to serrate;

stigma lobes pinkish to olive.

Fruits

indehiscent (rarely rupturing irregularly), scarlet or crimson, spheric to ovoid, 15–50 × 15–40 mm, fleshy, surfaces not hidden by widely spaced hairs in axils of scales;

scales 13–21, distal scales spine-tipped, minutely puberulent.

indehiscent or weakly dehiscent through basal abscission pore, pink or red, spheric to ovoid-cylindric, surfaces partly or entirely hidden by hairs from axils of scales and long areolar hairs of stem apex, usually quickly drying to tan shell before seed dispersal, 10–30 mm;

scales several, tips dark, spinelike, glabrous.

Seeds

black, spheric-reniform or irregularly obovoid, 2.5–3 mm, glossy;

testa cells flat or very slightly convex.

black or gray, angular or slightly wrinkled, spheric to obovoid, 2–3 mm;

testa cell surfaces slightly convex, with weak network pattern of slightly protruding anticlinal cell walls.

2n

= 22.

= 22.

Echinocactus texensis

Echinocactus horizonthalonius

Phenology Flowering late spring. Flowering Apr–Sep.
Habitat Chihuahuan Desert, grasslands, openings in oak woodlands, Tamaulipan thorn scrub, deep soils, saline flats, low limestone hills Arid rocky slopes, primarily limestone
Elevation 0-1400 m (0-4600 ft) 600-1700(-2500) m (2000-5600(-8200) ft)
Distribution
from FNA
NM; TX; Mexico (Coahuila, Durango, Nuevo León, Tamaulipas)
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM; TX; Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

The western, desert populations of Echinocactus texensis, unlike the eastern plants, have longer central spines that project stiffly outward and can flatten off-road vehicle tires or seriously injure a large mammal stepping on them. A dense cover of ephemeral herbs or shallow blanket of snow can hide this species completely from view.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The Sonoran Desert populations of Echinocactus horizonthalonius have been segregated as var. nicholii, but are relatively similar to plants in New Mexico and extreme western Texas. Much greater morphologic diversity exists farther east and in Mexico, where shorter-spined, nearly flat-topped plants, which are more distinctive than the Sonoran Desert populations, have escaped taxonomic distinction.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 4, p. 190. FNA vol. 4, p. 189.
Parent taxa Cactaceae > subfam. Cactoideae > Echinocactus Cactaceae > subfam. Cactoideae > Echinocactus
Sibling taxa
E. horizonthalonius, E. polycephalus
E. polycephalus, E. texensis
Synonyms Homalocephala texensis E. horizonthalonius var. nicholii
Name authority Hopffer: Allg. Gartenzeitung 10: 297. (1842) Lemaire: Cact. Gen. Sp. Nov., 19. (1839)
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