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keel wormseed

fetid glandular-goosefoot, fetid goosefoot

Stems

erect to ascending, branched, 3–5.5 dm, pilose with glandular trichomes or sessile glands.

erect, 2.3–5.2 dm, sparsely pubescent with short hairs, these rarely with gland heads.

Leaves

aromatic;

petiole 0.5–1.4 cm;

blade ovate to broadly ovate, 0.9–1.6 × 0.8–1.4 cm, somewhat reduced in inflorescence, base cuneate to truncate, apex acute, glandular-puberulose and pilosulose on veins.

aromatic;

petiole to 1.3 mm;

blade 1.7–4.5 × 0.7–2.6 cm, base cuneate, margins pinnatifid or entire (in distal leaves), apex acute to acuminate, mucronate, with sessile glands adaxially.

Inflorescences

axillary glomerules;

glomerules subglobose, 1.5–2.5 mm diam.;

bracts leaflike, ovate, 3–5 mm, margins crenate-dentate, apex acute.

terminal compound cymes, 8.5–22 cm;

bracts leaflike, lanceolate, 2.1–10.3 × 0.1–0.6 mm, generally absent in fruit.

Flowers

perianth segments 5, distinct nearly to base, distinct portions narrow-oblong, prominently keeled especially apically, 0.8–0.9 × 0.3–0.4 mm, apex acute, not crested, covered with septate hairs, becoming coriaceous and white in fruit;

stamens absent or 1;

stigmas 2.

perianth segments 5, distinct nearly to base, distinct portion ovate, with large single tubercle abaxially near apex, 0.5–0.7 × 0.35–0.6 mm, apex acuminate, with sessile golden glands, covering seed at maturity;

stamens 1(–2);

stigmas 2.

Achenes

ovoid;

pericarp adherent, membranaceous, slightly rugose.

subglobose;

pericarp adherent, membranaceous, papillose, becoming rugose, usually white-blotchy.

Seeds

reddish brown, ovoid, 0.6 × 0.3–0.4 mm, margins keeled over radicle and channeled over cotyledons;

seed coat smooth.

subglobose, 0.6–0.9 × 0.5–0.7 mm, margins rimmed;

seed coat rugose.

Dysphania carinata

Dysphania graveolens

Phenology Fruiting fall. Fruiting late summer–fall.
Habitat Waste areas on roadsides, sandy soils In shade of pines and junipers or occasionally in open dry plains, ridge tops, or in waste areas in the east
Elevation 10-100 m (0-300 ft) 900-2800 m (3000-9200 ft)
Distribution
from FNA
AL; AR; TX; native to Australia [Introduced in North America]
[BONAP county map]
from FNA
AZ; CO; LA; MA; ME; NM; NY; TX; UT; Mexico; Central America (Guatemala)
[WildflowerSearch map]
[BONAP county map]
Discussion

Western populations of Dysphania graveolens are native while those populations in the eastern part of the continent are considered introduced.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 4, p. 274. FNA vol. 4, p. 273.
Parent taxa Chenopodiaceae > Dysphania > sect. Orthospora Chenopodiaceae > Dysphania > sect. Botryoides > subsect. Incisa
Sibling taxa
D. ambrosioides, D. anthelmintica, D. aristata, D. botrys, D. chilensis, D. cristata, D. graveolens, D. multifida, D. pumilio
D. ambrosioides, D. anthelmintica, D. aristata, D. botrys, D. carinata, D. chilensis, D. cristata, D. multifida, D. pumilio
Synonyms Chenopodium carinatum Chenopodium graveolens, Chenopodium incisum, Teloxys graveolens
Name authority (R. Brown) Mosyakin & Clemants: Ukrayins’k. Bot. Zhurn., n. s. 59: 382. (2002) (Willdenow) Mosyakin & Clemants: in & Clemants, Ukrayins’k. Bot. Zhurn., n. s. 59: 383. (2002)
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