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keel wormseed

feather-geranium, Jerusalem oak goosefoot, Jerusalem-oak

Stems

erect to ascending, branched, 3–5.5 dm, pilose with glandular trichomes or sessile glands.

erect to ascending, branched at base to ± simple, 1–6(–10) dm, pubescent with short-stalked glandular hairs.

Leaves

aromatic;

petiole 0.5–1.4 cm;

blade ovate to broadly ovate, 0.9–1.6 × 0.8–1.4 cm, somewhat reduced in inflorescence, base cuneate to truncate, apex acute, glandular-puberulose and pilosulose on veins.

aromatic;

petiole to 2.5 mm;

blade 1.3–4 × 0.6–2.7 cm, base cuneate, margins lyrate-sinuate, pinnatifid, or occasionally entire (in distal leaves), apex acute to subobtuse, glandular-pubescent abaxially.

Inflorescences

axillary glomerules;

glomerules subglobose, 1.5–2.5 mm diam.;

bracts leaflike, ovate, 3–5 mm, margins crenate-dentate, apex acute.

axillary cymes, often arranged in terminal thyrses, 12–24 cm, subtended by cauline leaves;

bracts absent.

Flowers

perianth segments 5, distinct nearly to base, distinct portions narrow-oblong, prominently keeled especially apically, 0.8–0.9 × 0.3–0.4 mm, apex acute, not crested, covered with septate hairs, becoming coriaceous and white in fruit;

stamens absent or 1;

stigmas 2.

perianth segments 5, distinct nearly to base, distinct portion elliptic or ovate to oblong, 0.7–1.1 × 0.5–0.7 mm, apex acute to obtuse, rounded abaxially, densely glandular-pubescent, covering fruit at maturity;

stamens 1–3(–5);

stigmas 2.

Achenes

ovoid;

pericarp adherent, membranaceous, slightly rugose.

subglobose;

pericarp adherent, membranaceous, papillose, becoming rugose, usually white-blotchy.

Seeds

reddish brown, ovoid, 0.6 × 0.3–0.4 mm, margins keeled over radicle and channeled over cotyledons;

seed coat smooth.

globose to subglobose, (0.5–) 0.6–0.8 × 0.5–0.7 mm, margins rounded (rarely indistinctly furrowed);

seed coat rugose.

2n

= 18.

Dysphania carinata

Dysphania botrys

Phenology Fruiting fall. Fruiting Aug–Oct.
Habitat Waste areas on roadsides, sandy soils Sandy or gravelly soils, dry rocky ridges and cliffs, mud flats, waste places
Elevation 10-100 m (0-300 ft) 0-2000 m (0-6600 ft)
Distribution
from FNA
AL; AR; TX; native to Australia [Introduced in North America]
[BONAP county map]
from FNA
AZ; CA; CO; CT; DE; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OR; PA; RI; SD; TX; UT; VA; VT; WA; WI; WV; WY; BC; NB; NS; ON; PE; QC; s Europe; s Asia; c Asia; se Asia [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
Discussion

Dysphania botrys is related to a species from Africa and southern Eurasia, D. schraderiana (Schultes) Mosyakin & Clemants, which may occur locally in North America as introduced. Dysphania schraderiana has distinctly keeled perianth segments with mostly sessile or subsessile glands. The general inflorescence in D. schraderiana is usually leafy almost to the top, distal cauline leaves are similar to proximal ones (in D. botrys distal leaves are normally much reduced, and the distal portion of the general inflorescence appears nearly leafless). H. A. Wahl (1954) reported that D. schraderiana (as Chenopodium schraderianum) had been grown in Ontario. He did not indicate that it had escaped.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 4, p. 274. FNA vol. 4, p. 272.
Parent taxa Chenopodiaceae > Dysphania > sect. Orthospora Chenopodiaceae > Dysphania > sect. Botryoides > subsect. Botrys
Sibling taxa
D. ambrosioides, D. anthelmintica, D. aristata, D. botrys, D. chilensis, D. cristata, D. graveolens, D. multifida, D. pumilio
D. ambrosioides, D. anthelmintica, D. aristata, D. carinata, D. chilensis, D. cristata, D. graveolens, D. multifida, D. pumilio
Synonyms Chenopodium carinatum Chenopodium botrys, Teloxys botrys
Name authority (R. Brown) Mosyakin & Clemants: Ukrayins’k. Bot. Zhurn., n. s. 59: 382. (2002) (Linnaeus) Mosyakin & Clemants: Ukrayins’k. Bot. Zhurn., n. s. 59: 383. (2002)
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