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feather-geranium, Jerusalem oak goosefoot, Jerusalem-oak

wormseed

Stems

erect to ascending, branched at base to ± simple, 1–6(–10) dm, pubescent with short-stalked glandular hairs.

erect, [very bushy-branched] profusely branched from base to ± simple, [0.4–]0.7–2.5[–5] dm, glabrous or with scattered inflated hairs and uniseriate nonglandular trichomes, especially towards base.

Leaves

aromatic;

petiole to 2.5 mm;

blade 1.3–4 × 0.6–2.7 cm, base cuneate, margins lyrate-sinuate, pinnatifid, or occasionally entire (in distal leaves), apex acute to subobtuse, glandular-pubescent abaxially.

non-aromatic;

petiole to 0.4 mm;

blade [0.8–]2.8–3.4[–4.1] × 0.1–0.3[–0.6] cm, base attenuate, margins entire (to occasionally shallowly erose-dentate), apex acute, mucronate, glabrous adaxially.

Inflorescences

axillary cymes, often arranged in terminal thyrses, 12–24 cm, subtended by cauline leaves;

bracts absent.

a terminal thyrse, 2.2–4.5[–13] cm, or lax, pyramidal cymes, bearing flowers almost from base;

bracts absent.

Flowers

perianth segments 5, distinct nearly to base, distinct portion elliptic or ovate to oblong, 0.7–1.1 × 0.5–0.7 mm, apex acute to obtuse, rounded abaxially, densely glandular-pubescent, covering fruit at maturity;

stamens 1–3(–5);

stigmas 2.

perianth segments 5, distinct nearly to base, distinct portions obovate to elliptic or ovate, with low tubercle or sometimes keeled abaxially, 0.5–0.7 × 0.3–0.4 mm, apex obtuse to subacute, glabrous, loosely covering fruit at maturity;

stamens 5 (or absent in pistillate flowers in distal portion of inflorescence);

stigmas 2.

Achenes

subglobose;

pericarp adherent, membranaceous, papillose, becoming rugose, usually white-blotchy.

subglobose;

pericarp adherent, membranaceous, finely granular.

Seeds

globose to subglobose, (0.5–) 0.6–0.8 × 0.5–0.7 mm, margins rounded (rarely indistinctly furrowed);

seed coat rugose.

subglobose, 0.5–0.8 × 0.4–0.5 mm, margins rimmed;

seed coat smooth.

2n

= 18.

Dysphania botrys

Dysphania aristata

Phenology Fruiting Aug–Oct. Fruiting late summer–fall.
Habitat Sandy or gravelly soils, dry rocky ridges and cliffs, mud flats, waste places Waste areas, sandy soils
Elevation 0-2000 m (0-6600 ft) 0-500 m (0-1600 ft)
Distribution
from FNA
AZ; CA; CO; CT; DE; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OR; PA; RI; SD; TX; UT; VA; VT; WA; WI; WV; WY; BC; NB; NS; ON; PE; QC; s Europe; s Asia; c Asia; se Asia [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
MI; NY; native to Asia [Introduced in North America; introduced in s, se Europe]
[BONAP county map]
Discussion

Dysphania botrys is related to a species from Africa and southern Eurasia, D. schraderiana (Schultes) Mosyakin & Clemants, which may occur locally in North America as introduced. Dysphania schraderiana has distinctly keeled perianth segments with mostly sessile or subsessile glands. The general inflorescence in D. schraderiana is usually leafy almost to the top, distal cauline leaves are similar to proximal ones (in D. botrys distal leaves are normally much reduced, and the distal portion of the general inflorescence appears nearly leafless). H. A. Wahl (1954) reported that D. schraderiana (as Chenopodium schraderianum) had been grown in Ontario. He did not indicate that it had escaped.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 4, p. 272. FNA vol. 4, p. 273.
Parent taxa Chenopodiaceae > Dysphania > sect. Botryoides > subsect. Botrys Chenopodiaceae > Dysphania > sect. Botryoides > subsect. Teloxys
Sibling taxa
D. ambrosioides, D. anthelmintica, D. aristata, D. carinata, D. chilensis, D. cristata, D. graveolens, D. multifida, D. pumilio
D. ambrosioides, D. anthelmintica, D. botrys, D. carinata, D. chilensis, D. cristata, D. graveolens, D. multifida, D. pumilio
Synonyms Chenopodium botrys, Teloxys botrys Chenopodium aristatum
Name authority (Linnaeus) Mosyakin & Clemants: Ukrayins’k. Bot. Zhurn., n. s. 59: 383. (2002) (Linnaeus) Mosyakin & Clemants: Ukrayins’k. Bot. Zhurn., n. s. 59: 383. (2002)
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