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southern wood fern

dryoptère de Goldie, Goldie's wood fern

Leaves

somewhat dimorphic, green through winter, 35–120 × 10–30 cm.

monomorphic, dying back in winter, 35–120 × 15–40 cm.

Petiole

more than 1/4 length of leaf, scaly at base;

scales scattered, brown.

1/3 length of leaf, scaly at base;

scales scattered, dark, glossy brown to nearly black, with pale border.

Blade

dark green, lanceolate, pinnate-pinnatifid, herbaceous, not glandular.

green, often white-mottled at tip, ovate, tapering abruptly at apex, pinnate-pinnatifid to 2-pinnate at base, herbaceous, not glandular.

Pinnae

nearly in plane of blade, lance-oblong;

fertile pinnae in distal 1/2 of leaf, distinctly narrower than proximal vegetative pinnae (only in this species);

basal pinnae lanceolate-oblong, much reduced, basal pinnules slightly shorter than adjacent pinnules, basal basiscopic pinnule slightly longer than basal acroscopic pinnule;

pinnule margins distantly serrate.

parallel to plane of blade, ovate-lanceolate, broadest above base;

basal pinnae broadly oblong-lanceolate, slightly reduced, basal pinnule equal to adjacent pinnules, basal basiscopic pinnule and basal acroscopic pinnule equal;

pinnule margins crenulate or serrate.

Indusia

lacking glands.

and axes lacking glands.

Sori

midway between midvein and margin of segments.

nearer midvein than margin.

2n

= 82.

= 82.

Dryopteris ludoviciana

Dryopteris goldieana

Habitat Swamps and wet woods Dense, moist woods, especially ravines, limey seeps, or at the edge of swamps
Elevation 0–100 m (0–300 ft) 50–1500 m (200–4900 ft)
Distribution
from FNA
AL; AR; FL; GA; LA; NC; SC
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; CT; DE; GA; IA; IL; IN; KY; MA; MD; ME; MI; MN; MO; NC; NH; NJ; NY; OH; PA; RI; TN; VA; VT; WI; WV; NB; ON; QC
[WildflowerSearch map]
Discussion

Dryopteris ludoviciana is endemic to southeastern United States. This diploid is one of the parents of D. celsa and D. cristata. It crosses with D. celsa to produce sterile hybrids.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Dryopteris goldieana is diploid and is one of the parents of D. celsa and of D. clintoniana. Dryopteris goldieana hybridizes with five species. Hybrids can be identified by the glossy dark scales and large blade size. A remarkable additional hybrid (× Dryostichum singulare W. H. Wagner), involving this species and Polystichum lonchitis, is known from Gray and Simcoe counties, Ontario. It is intermediate between the parents and is sterile (W. H. Wagner Jr., F. S. Wagner et al. 1992).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Dryopteridaceae > Dryopteris Dryopteridaceae > Dryopteris
Sibling taxa
D. arguta, D. campyloptera, D. carthusiana, D. celsa, D. cinnamomea, D. clintoniana, D. cristata, D. expansa, D. filix-mas, D. fragrans, D. goldieana, D. intermedia, D. marginalis
D. arguta, D. campyloptera, D. carthusiana, D. celsa, D. cinnamomea, D. clintoniana, D. cristata, D. expansa, D. filix-mas, D. fragrans, D. intermedia, D. ludoviciana, D. marginalis
Synonyms Aspidium ludovicianum, D. floridana Aspidium goldieanum
Name authority (Kunze) Small: Ferns S. E. States 281. (1938) (Hooker ex Goldie) A. Gray: Manual 631. (1848) — (as goldiana)
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