Dryopteris goldieana |
Dryopteris clintoniana |
|
---|---|---|
dryoptère de Goldie, Goldie's wood fern |
Clinton's wood fern, dryoptère de Clinton |
|
Leaves | monomorphic, dying back in winter, 35–120 × 15–40 cm. |
dimorphic, 45–100 × 12–20 cm; fertile leaves dying back in winter; sterile leaves 1–several, smaller, green through winter. |
Petiole | 1/3 length of leaf, scaly at base; scales scattered, dark, glossy brown to nearly black, with pale border. |
1/4–1/3 length of leaf, scaly at least at base; scales scattered, tan, sometimes with dark brown center. |
Blade | green, often white-mottled at tip, ovate, tapering abruptly at apex, pinnate-pinnatifid to 2-pinnate at base, herbaceous, not glandular. |
green, lanceolate, with nearly parallel sides, pinnate-pinnatifid, herbaceous, not glandular. |
Pinnae | parallel to plane of blade, ovate-lanceolate, broadest above base; basal pinnae broadly oblong-lanceolate, slightly reduced, basal pinnule equal to adjacent pinnules, basal basiscopic pinnule and basal acroscopic pinnule equal; pinnule margins crenulate or serrate. |
of fertile leaves twisted out of plane of blade but not fully perpendicular to it, narrowly elongate-deltate; basal pinnae narrowly elongate-deltate, much reduced; basal pinnules longer than or equal to adjacent pinnules, basal basiscopic pinnule and basal acroscopic pinnule equal; pinnule margins serrate or biserrate, with spiny teeth. |
Indusia | and axes lacking glands. |
lacking glands. |
Sori | nearer midvein than margin. |
midway between midvein and margin of segments. |
2n | = 82. |
= 246. |
Dryopteris goldieana |
Dryopteris clintoniana |
|
Habitat | Dense, moist woods, especially ravines, limey seeps, or at the edge of swamps | Swampy woods |
Elevation | 50–1500 m (200–4900 ft) | 50–600 m (200–2000 ft) |
Distribution |
AL; CT; DE; GA; IA; IL; IN; KY; MA; MD; ME; MI; MN; MO; NC; NH; NJ; NY; OH; PA; RI; TN; VA; VT; WI; WV; NB; ON; QC
|
CT; IN; MA; ME; MI; NH; NJ; NY; OH; PA; RI; VT; NB; ON; QC
|
Discussion | Dryopteris goldieana is diploid and is one of the parents of D. celsa and of D. clintoniana. Dryopteris goldieana hybridizes with five species. Hybrids can be identified by the glossy dark scales and large blade size. A remarkable additional hybrid (× Dryostichum singulare W. H. Wagner), involving this species and Polystichum lonchitis, is known from Gray and Simcoe counties, Ontario. It is intermediate between the parents and is sterile (W. H. Wagner Jr., F. S. Wagner et al. 1992). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Dryopteris clintoniana is a North American endemic and an allohexaploid derived from D. cristata and D. goldieana. Dryopteris clintoniana hybridizes with six species. Hybrids can be identified by the fairly narrow blades and elongate-deltate proximal pinnae. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Dryopteridaceae > Dryopteris | Dryopteridaceae > Dryopteris |
Sibling taxa | ||
Synonyms | Aspidium goldieanum | Aspidium cristatum var. clintonianum, D. cristata var. clintoniana |
Name authority | (Hooker ex Goldie) A. Gray: Manual 631. (1848) — (as goldiana) | (D. C. Eaton) Dowell: Proc. Staten Island Assoc. Arts 1: 64. (1906) |
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