Dryopteris fragrans |
Dryopteris filix-mas |
|
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dryoptère fragrante, fragrant wood fern |
dryoptère fougère mâle, male fern, male wood fern |
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Leaves | monomorphic, green through winter, 6–40 × 1–6 cm, old leaves persistent as gray or brown conspicuous clump at plant base (only in this species). |
monomorphic, dying back in winter, 28–120 × 10–30 cm. |
Petiole | 1/3 length of leaf, scaly throughout; scales dense, brown to red-brown. |
less than 1/4 length of leaf, scaly at least at base; scales scattered, brown, of 2 distinct kinds, 1 broad, 1 hairlike (only this species has 2 distinct forms of scales without intermediates). |
Blade(s) | green, linear-lanceolate, pinnate-pinnatifid to 2-pinnate, papery, densely scaly (only in this species) abaxially, glandular, aromatic when handled (only in this species). |
green, ovate-lanceolate, pinnate-pinnatifid to 2-pinnate at base, firm but not leathery, not glandular. |
Pinnae | ± in plane of blade, linear-oblong; basal pinnae linear-oblong, much reduced, basal pinnules longer than adjacent pinnules, basal basiscopic pinnule and basal acroscopic pinnule equal; pinnule margins crenately toothed. |
± in plane of blade, lanceolate; basal pinnae ovate-lanceolate, much reduced, basal pinnules or segments ± same length as adjacent pinnules, basal basiscopic pinnule and basal acroscopic pinnule equal; pinnule margins serrate to lobed. |
Indusia | glandular. |
lacking glands. |
Sori | midway between midvein and margin of segments. |
midway between midvein and margin of segments. |
2n | = 82. |
= 164. |
Dryopteris fragrans |
Dryopteris filix-mas |
|
Habitat | Shaded cliffs and talus, often of limestone | Dense woods and talus slopes on limestone (ne North America), open woods among boulders and talus of granite or igneous rock (Rocky Mountains) |
Elevation | 50–1800 m (200–5900 ft) | 200–2500 m (700–8200 ft) |
Distribution |
AK; ME; MI; MN; NH; NY; VT; WI; AB; BC; MB; NB; NF; NS; NT; ON; QC; SK; YT; Greenland; Europe in n Finland; Asia in n, nw former Soviet republics
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AZ; CA; CO; ID; MI; MT; NM; NV; OK; OR; TX; UT; WA; WI; WY; AB; BC; NF; NS; ON; QC; SK; Greenland; Europe; Asia
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Discussion | Dryopteris fragrans is a northern species and is not closely related to the other species in North America. The only hybrid known to involve D. fragrans is with D. marginalis, producing D. × algonquinensis D. Britton. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The taxonomy of Dryopteris filix-mas is not well understood. In North America, this fern has been considered both an auto- and an allopolyploid and may be composed of at least two closely related taxa. Plants in the northeast and northwest are tetraploid. These differ morphologically and ecologically from a taxon of unknown chromosome number in the southwestern Rocky Mountains. The Rocky Mountain taxon closely resembles the Mexican D. pseudofilix-mas (Fée) Rothmaler. Dryopteris filix-mas also occurs in Europe, and it is known to be an allopolyploid of D. caucasica (A. Braun) Fraser-Jenkins & Corley × oreades Fomin. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Dryopteridaceae > Dryopteris | Dryopteridaceae > Dryopteris |
Sibling taxa | ||
Synonyms | Polypodium fragrans, D. fragrans var. remotiuscula | Polypodium filix-mas |
Name authority | (Linnaeus) Schott: Gen. Fil. plate 9. (1834) | (Linnaeus) Schott: Gen. Fil. plate 67. (1834) |
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