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dryoptère fragrante, fragrant wood fern

dryoptère fougère mâle, male fern, male wood fern

Leaves

monomorphic, green through winter, 6–40 × 1–6 cm, old leaves persistent as gray or brown conspicuous clump at plant base (only in this species).

monomorphic, dying back in winter, 28–120 × 10–30 cm.

Petiole

1/3 length of leaf, scaly throughout;

scales dense, brown to red-brown.

less than 1/4 length of leaf, scaly at least at base;

scales scattered, brown, of 2 distinct kinds, 1 broad, 1 hairlike (only this species has 2 distinct forms of scales without intermediates).

Blade(s)

green, linear-lanceolate, pinnate-pinnatifid to 2-pinnate, papery, densely scaly (only in this species) abaxially, glandular, aromatic when handled (only in this species).

green, ovate-lanceolate, pinnate-pinnatifid to 2-pinnate at base, firm but not leathery, not glandular.

Pinnae

± in plane of blade, linear-oblong;

basal pinnae linear-oblong, much reduced, basal pinnules longer than adjacent pinnules, basal basiscopic pinnule and basal acroscopic pinnule equal;

pinnule margins crenately toothed.

± in plane of blade, lanceolate;

basal pinnae ovate-lanceolate, much reduced, basal pinnules or segments ± same length as adjacent pinnules, basal basiscopic pinnule and basal acroscopic pinnule equal;

pinnule margins serrate to lobed.

Indusia

glandular.

lacking glands.

Sori

midway between midvein and margin of segments.

midway between midvein and margin of segments.

2n

= 82.

= 164.

Dryopteris fragrans

Dryopteris filix-mas

Habitat Shaded cliffs and talus, often of limestone Dense woods and talus slopes on limestone (ne North America), open woods among boulders and talus of granite or igneous rock (Rocky Mountains)
Elevation 50–1800 m (200–5900 ft) 200–2500 m (700–8200 ft)
Distribution
from FNA
AK; ME; MI; MN; NH; NY; VT; WI; AB; BC; MB; NB; NF; NS; NT; ON; QC; SK; YT; Greenland; Europe in n Finland; Asia in n, nw former Soviet republics
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; ID; MI; MT; NM; NV; OK; OR; TX; UT; WA; WI; WY; AB; BC; NF; NS; ON; QC; SK; Greenland; Europe; Asia
[WildflowerSearch map]
[BONAP county map]
Discussion

Dryopteris fragrans is a northern species and is not closely related to the other species in North America. The only hybrid known to involve D. fragrans is with D. marginalis, producing D. × algonquinensis D. Britton.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The taxonomy of Dryopteris filix-mas is not well understood. In North America, this fern has been considered both an auto- and an allopolyploid and may be composed of at least two closely related taxa. Plants in the northeast and northwest are tetraploid. These differ morphologically and ecologically from a taxon of unknown chromosome number in the southwestern Rocky Mountains. The Rocky Mountain taxon closely resembles the Mexican D. pseudofilix-mas (Fée) Rothmaler. Dryopteris filix-mas also occurs in Europe, and it is known to be an allopolyploid of D. caucasica (A. Braun) Fraser-Jenkins & Corley × oreades Fomin.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Dryopteridaceae > Dryopteris Dryopteridaceae > Dryopteris
Sibling taxa
D. arguta, D. campyloptera, D. carthusiana, D. celsa, D. cinnamomea, D. clintoniana, D. cristata, D. expansa, D. filix-mas, D. goldieana, D. intermedia, D. ludoviciana, D. marginalis
D. arguta, D. campyloptera, D. carthusiana, D. celsa, D. cinnamomea, D. clintoniana, D. cristata, D. expansa, D. fragrans, D. goldieana, D. intermedia, D. ludoviciana, D. marginalis
Synonyms Polypodium fragrans, D. fragrans var. remotiuscula Polypodium filix-mas
Name authority (Linnaeus) Schott: Gen. Fil. plate 9. (1834) (Linnaeus) Schott: Gen. Fil. plate 67. (1834)
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