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log fern

dryoptère de Goldie, Goldie's wood fern

Leaves

monomorphic, dying back in winter, 65–120 × 15–30 cm.

monomorphic, dying back in winter, 35–120 × 15–40 cm.

Petiole

1/3 length of leaf, scaly at least at base;

scales scattered, dark brown or tan with dark central stripe.

1/3 length of leaf, scaly at base;

scales scattered, dark, glossy brown to nearly black, with pale border.

Blade

green, ovate-lanceolate, gradually tapering to tip, pinnate-pinnatifid, herbaceous, not glandular.

green, often white-mottled at tip, ovate, tapering abruptly at apex, pinnate-pinnatifid to 2-pinnate at base, herbaceous, not glandular.

Pinnae

± in plane of blade, lanceolate-ovate;

basal pinnae linear-oblong, much reduced, basal pinnules longer than adjacent pinnules, basal basiscopic pinnule and basal acroscopic pinnule equal;

pinnule margins crenately toothed.

parallel to plane of blade, ovate-lanceolate, broadest above base;

basal pinnae broadly oblong-lanceolate, slightly reduced, basal pinnule equal to adjacent pinnules, basal basiscopic pinnule and basal acroscopic pinnule equal;

pinnule margins crenulate or serrate.

Indusia

lacking glands.

and axes lacking glands.

Sori

midway between midvein and margin of segments.

nearer midvein than margin.

2n

= 164.

= 82.

Dryopteris celsa

Dryopteris goldieana

Habitat Seepage slopes, hammocks and logs in swamps, mostly on the Piedmont and Coastal Plain Dense, moist woods, especially ravines, limey seeps, or at the edge of swamps
Elevation 50–800 m (200–2600 ft) 50–1500 m (200–4900 ft)
Distribution
from FNA
AL; AR; DE; GA; IL; KY; LA; MD; MI; MO; NC; NJ; NY; PA; SC; TN; VA; WV
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; CT; DE; GA; IA; IL; IN; KY; MA; MD; ME; MI; MN; MO; NC; NH; NJ; NY; OH; PA; RI; TN; VA; VT; WI; WV; NB; ON; QC
[WildflowerSearch map]
Discussion

Dryopteris celsa is a fertile allotetraploid derived from hybridization between D. goldieana and D. ludoviciana. Dryopteris celsa hybridizes with six species; hybrids can usually be identified by the dark-striped scales.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Dryopteris goldieana is diploid and is one of the parents of D. celsa and of D. clintoniana. Dryopteris goldieana hybridizes with five species. Hybrids can be identified by the glossy dark scales and large blade size. A remarkable additional hybrid (× Dryostichum singulare W. H. Wagner), involving this species and Polystichum lonchitis, is known from Gray and Simcoe counties, Ontario. It is intermediate between the parents and is sterile (W. H. Wagner Jr., F. S. Wagner et al. 1992).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Dryopteridaceae > Dryopteris Dryopteridaceae > Dryopteris
Sibling taxa
D. arguta, D. campyloptera, D. carthusiana, D. cinnamomea, D. clintoniana, D. cristata, D. expansa, D. filix-mas, D. fragrans, D. goldieana, D. intermedia, D. ludoviciana, D. marginalis
D. arguta, D. campyloptera, D. carthusiana, D. celsa, D. cinnamomea, D. clintoniana, D. cristata, D. expansa, D. filix-mas, D. fragrans, D. intermedia, D. ludoviciana, D. marginalis
Synonyms D. goldiana subsp. celsa Aspidium goldieanum
Name authority (W. Palmer) Knowlton: W. Palmer, & Pollard, Proc. Biol. Soc. Wash. 13: 202. (1900) (Hooker ex Goldie) A. Gray: Manual 631. (1848) — (as goldiana)
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