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log fern

crested wood fern, dryoptère à crêtes, shield fern

Leaves

monomorphic, dying back in winter, 65–120 × 15–30 cm.

dimorphic, 35–70 × 8–12 cm;

fertile leaves dying back in winter;

sterile leaves several, small, green through winter, forming "rosette." Petiole 1/4–1/3 length of leaf, scaly at least at base;

scales scattered, tan.

Petiole

1/3 length of leaf, scaly at least at base;

scales scattered, dark brown or tan with dark central stripe.

Blade

green, ovate-lanceolate, gradually tapering to tip, pinnate-pinnatifid, herbaceous, not glandular.

green, narrowly lanceolate or with parallel sides, pinnate-pinnatifid, not glandular.

Pinnae

± in plane of blade, lanceolate-ovate;

basal pinnae linear-oblong, much reduced, basal pinnules longer than adjacent pinnules, basal basiscopic pinnule and basal acroscopic pinnule equal;

pinnule margins crenately toothed.

of fertile leaves twisted out of plane of blade and perpendicular to it, deltate;

basal pinnae deltate, somewhat reduced, basal pinnules longer than adjacent pinnules, basal basiscopic pinnule and basal acroscopic pinnule equal;

pinnule margins distantly serrate, with spiny teeth.

Indusia

lacking glands.

lacking glands.

Sori

midway between midvein and margin of segments.

midway between midvein and margin of segments.

2n

= 164.

= 164.

Dryopteris celsa

Dryopteris cristata

Habitat Seepage slopes, hammocks and logs in swamps, mostly on the Piedmont and Coastal Plain Swamps, swampy woods, or open shrubby wetlands
Elevation 50–800 m (200–2600 ft) 0–1200 m (0–3900 ft)
Distribution
from FNA
AL; AR; DE; GA; IL; KY; LA; MD; MI; MO; NC; NJ; NY; PA; SC; TN; VA; WV
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; CT; DE; IA; IL; IN; MA; MD; ME; MI; MN; MT; NC; ND; NE; NH; NJ; NY; OH; PA; RI; TN; VA; VT; WI; WV; AB; BC; MB; NB; NF; NS; ON; PE; QC; SK; Europe
[WildflowerSearch map]
[BONAP county map]
Discussion

Dryopteris celsa is a fertile allotetraploid derived from hybridization between D. goldieana and D. ludoviciana. Dryopteris celsa hybridizes with six species; hybrids can usually be identified by the dark-striped scales.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Dryopteris cristata is believed to be an allotetraploid derived from D. ludoviciana and an unknown diploid called " D. semicristata " by W. H. Wagner Jr. (1971). This ancestral taxon could have been either North American or Eurasian and may have become extinct during the last glaciation (T. J. Carlson and W. H. Wagner Jr. 1982). Dryopteris cristata hybridizes with five species; these hybrids can be identified by the narrow blades and deltate proximal pinnae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Dryopteridaceae > Dryopteris Dryopteridaceae > Dryopteris
Sibling taxa
D. arguta, D. campyloptera, D. carthusiana, D. cinnamomea, D. clintoniana, D. cristata, D. expansa, D. filix-mas, D. fragrans, D. goldieana, D. intermedia, D. ludoviciana, D. marginalis
D. arguta, D. campyloptera, D. carthusiana, D. celsa, D. cinnamomea, D. clintoniana, D. expansa, D. filix-mas, D. fragrans, D. goldieana, D. intermedia, D. ludoviciana, D. marginalis
Synonyms D. goldiana subsp. celsa Polypodium cristatum
Name authority (W. Palmer) Knowlton: W. Palmer, & Pollard, Proc. Biol. Soc. Wash. 13: 202. (1900) (Linnaeus) A. Gray: Manual 631. (1848)
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