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log fern

Clinton's wood fern, dryoptère de Clinton

Leaves

monomorphic, dying back in winter, 65–120 × 15–30 cm.

dimorphic, 45–100 × 12–20 cm;

fertile leaves dying back in winter;

sterile leaves 1–several, smaller, green through winter.

Petiole

1/3 length of leaf, scaly at least at base;

scales scattered, dark brown or tan with dark central stripe.

1/4–1/3 length of leaf, scaly at least at base;

scales scattered, tan, sometimes with dark brown center.

Blade

green, ovate-lanceolate, gradually tapering to tip, pinnate-pinnatifid, herbaceous, not glandular.

green, lanceolate, with nearly parallel sides, pinnate-pinnatifid, herbaceous, not glandular.

Pinnae

± in plane of blade, lanceolate-ovate;

basal pinnae linear-oblong, much reduced, basal pinnules longer than adjacent pinnules, basal basiscopic pinnule and basal acroscopic pinnule equal;

pinnule margins crenately toothed.

of fertile leaves twisted out of plane of blade but not fully perpendicular to it, narrowly elongate-deltate;

basal pinnae narrowly elongate-deltate, much reduced;

basal pinnules longer than or equal to adjacent pinnules, basal basiscopic pinnule and basal acroscopic pinnule equal;

pinnule margins serrate or biserrate, with spiny teeth.

Indusia

lacking glands.

lacking glands.

Sori

midway between midvein and margin of segments.

midway between midvein and margin of segments.

2n

= 164.

= 246.

Dryopteris celsa

Dryopteris clintoniana

Habitat Seepage slopes, hammocks and logs in swamps, mostly on the Piedmont and Coastal Plain Swampy woods
Elevation 50–800 m (200–2600 ft) 50–600 m (200–2000 ft)
Distribution
from FNA
AL; AR; DE; GA; IL; KY; LA; MD; MI; MO; NC; NJ; NY; PA; SC; TN; VA; WV
[WildflowerSearch map]
[BONAP county map]
from FNA
CT; IN; MA; ME; MI; NH; NJ; NY; OH; PA; RI; VT; NB; ON; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Dryopteris celsa is a fertile allotetraploid derived from hybridization between D. goldieana and D. ludoviciana. Dryopteris celsa hybridizes with six species; hybrids can usually be identified by the dark-striped scales.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Dryopteris clintoniana is a North American endemic and an allohexaploid derived from D. cristata and D. goldieana. Dryopteris clintoniana hybridizes with six species. Hybrids can be identified by the fairly narrow blades and elongate-deltate proximal pinnae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Dryopteridaceae > Dryopteris Dryopteridaceae > Dryopteris
Sibling taxa
D. arguta, D. campyloptera, D. carthusiana, D. cinnamomea, D. clintoniana, D. cristata, D. expansa, D. filix-mas, D. fragrans, D. goldieana, D. intermedia, D. ludoviciana, D. marginalis
D. arguta, D. campyloptera, D. carthusiana, D. celsa, D. cinnamomea, D. cristata, D. expansa, D. filix-mas, D. fragrans, D. goldieana, D. intermedia, D. ludoviciana, D. marginalis
Synonyms D. goldiana subsp. celsa Aspidium cristatum var. clintonianum, D. cristata var. clintoniana
Name authority (W. Palmer) Knowlton: W. Palmer, & Pollard, Proc. Biol. Soc. Wash. 13: 202. (1900) (D. C. Eaton) Dowell: Proc. Staten Island Assoc. Arts 1: 64. (1906)
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