Flora of North America species comparison

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Dryopteris celsa

Dryopteris carthusiana

log fern

dryoptère de cartheuser, spinulose fern, spinulose wood fern, tooth wood fern
Leaves

monomorphic, dying back in winter, 65–120 × 15–30 cm.

monomorphic, dying in winter, 15–75 × 10–30 cm.
Petiole

1/3 length of leaf, scaly at least at base;

scales scattered, dark brown or tan with dark central stripe.

1/4–1/3 length of leaf, scaly at least at base;

scales scattered, tan.
Blade

green, ovate-lanceolate, gradually tapering to tip, pinnate-pinnatifid, herbaceous, not glandular.

light green, ovate-lanceolate, 2–3-pinnate-pinnatifid, herbaceous, not glandular.
Pinnae

± in plane of blade, lanceolate-ovate;

basal pinnae linear-oblong, much reduced, basal pinnules longer than adjacent pinnules, basal basiscopic pinnule and basal acroscopic pinnule equal;

pinnule margins crenately toothed.

± in plane of blade, lance-oblong;

basal pinnae lanceolate-deltate, slightly reduced, basal pinnules usually longer than adjacent pinnules, basal basiscopic pinnule longer than basal acroscopic pinnule;

pinnule margins serrate, teeth spiny.
Indusia

lacking glands.

lacking glands.
Sori

midway between midvein and margin of segments.

midway between midvein and margin of segments.
2n

= 164.

= 164.

Dryopteris celsa

Dryopteris carthusiana
Habitat Seepage slopes, hammocks and logs in swamps, mostly on the Piedmont and Coastal Plain Swampy woods, moist wooded slopes, stream banks, and conifer plantations
Elevation 50–800 m (200–2600 ft) 0–1200 m (0–3900 ft)
Distribution
from FNA
AL; AR; DE; GA; IL; KY; LA; MD; MI; MO; NC; NJ; NY; PA; SC; TN; VA; WV
[WildflowerSearch map]
[BONAP county map]
from FNA
AR; CT; DE; IA; ID; IL; IN; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NY; OH; PA; RI; SC; TN; VA; VT; WA; WI; WV; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; YT; Eurasia
[WildflowerSearch map]
[BONAP county map]
Discussion

Dryopteris celsa is a fertile allotetraploid derived from hybridization between D. goldieana and D. ludoviciana. Dryopteris celsa hybridizes with six species; hybrids can usually be identified by the dark-striped scales.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Dryopteris carthusiana is tetraploid. Dryopteris intermedia is one parent, as indicated by chromosome pairing in their hybrid D. × triploidea Wherry. The other parent is the hypothetical missing ancestral species " D. semicristata " (see discussion for D. cristata). Dryopteris carthusiana hybridizes with five species; hybrids can be separated from D. intermedia by the lack of glandular hairs and by having 2-pinnate leaves.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Source FNA vol. 2. FNA vol. 2.
Parent taxa Dryopteridaceae > Dryopteris Dryopteridaceae > Dryopteris
Sibling taxa
D. arguta, D. campyloptera, D. carthusiana, D. cinnamomea, D. clintoniana, D. cristata, D. expansa, D. filix-mas, D. fragrans, D. goldieana, D. intermedia, D. ludoviciana, D. marginalis
D. arguta, D. campyloptera, D. celsa, D. cinnamomea, D. clintoniana, D. cristata, D. expansa, D. filix-mas, D. fragrans, D. goldieana, D. intermedia, D. ludoviciana, D. marginalis
Synonyms D. goldiana subsp. celsa Polypodium carthusianum, D. austriaca var. spinulosa, D. spinulosa, Polypodium spinulosum
Name authority (W. Palmer) Knowlton: W. Palmer, & Pollard, Proc. Biol. Soc. Wash. 13: 202. (1900) (Villars) H. P. Fuchs: Bull. Soc. Bot. France 105: 339. (1959)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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