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log fern

California wood fern, coastal fern, coastal wood fern, marginal wood fern, wood fern

Leaves

monomorphic, dying back in winter, 65–120 × 15–30 cm.

monomorphic, green through winter, 25–90 × 8–30 cm.

Petiole

1/3 length of leaf, scaly at least at base;

scales scattered, dark brown or tan with dark central stripe.

1/4–1/3 length of leaf, scaly at least at base;

scales scattered, light brown.

Blade

green, ovate-lanceolate, gradually tapering to tip, pinnate-pinnatifid, herbaceous, not glandular.

green to yellow-green, ovate-lanceolate, pinnate-pinnatifid to 2-pinnate at base, herbaceous, glandular.

Pinnae

± in plane of blade, lanceolate-ovate;

basal pinnae linear-oblong, much reduced, basal pinnules longer than adjacent pinnules, basal basiscopic pinnule and basal acroscopic pinnule equal;

pinnule margins crenately toothed.

± in plane of blade, lance-oblong;

basal pinnae deltate, not much reduced, basal pinnules ± same length as adjacent pinnules, basal basiscopic pinnule and basal acroscopic pinnule ± equal, pinnule margins serrate with spreading, spinelike teeth.

Indusia

lacking glands.

lacking glands.

Sori

midway between midvein and margin of segments.

midway between midvein and margin of segments.

2n

= 164.

= 82.

Dryopteris celsa

Dryopteris arguta

Habitat Seepage slopes, hammocks and logs in swamps, mostly on the Piedmont and Coastal Plain Shaded slopes and open woods
Elevation 50–800 m (200–2600 ft) 0–2100 m (0–6900 ft)
Distribution
from FNA
AL; AR; DE; GA; IL; KY; LA; MD; MI; MO; NC; NJ; NY; PA; SC; TN; VA; WV
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; OR; WA; BC; Mexico in Baja California
[WildflowerSearch map]
[BONAP county map]
Discussion

Dryopteris celsa is a fertile allotetraploid derived from hybridization between D. goldieana and D. ludoviciana. Dryopteris celsa hybridizes with six species; hybrids can usually be identified by the dark-striped scales.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Dryopteris arguta is somewhat variable. It has been suggested that more than one taxon is involved. No hybrids involving D. arguta are known.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Dryopteridaceae > Dryopteris Dryopteridaceae > Dryopteris
Sibling taxa
D. arguta, D. campyloptera, D. carthusiana, D. cinnamomea, D. clintoniana, D. cristata, D. expansa, D. filix-mas, D. fragrans, D. goldieana, D. intermedia, D. ludoviciana, D. marginalis
D. campyloptera, D. carthusiana, D. celsa, D. cinnamomea, D. clintoniana, D. cristata, D. expansa, D. filix-mas, D. fragrans, D. goldieana, D. intermedia, D. ludoviciana, D. marginalis
Synonyms D. goldiana subsp. celsa Aspidium argutum
Name authority (W. Palmer) Knowlton: W. Palmer, & Pollard, Proc. Biol. Soc. Wash. 13: 202. (1900) (Kaulfuss) Maxon: Amer. Fern J. 11: 3. (1921)
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