Flora of North America species comparison

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Drymocallis pseudorupestris

Drymocallis pseudorupestris var. saxicola
Drymocallis pseudorupestris

cliff drymocallis, cliff woodbeauty, false rock loving cinquefoil, Rocky Mountain sticky cinquefoil
Photo is of parent taxon

cliff drymocallis or wood beauty, cliff woodbeauty, Rocky Mountain sticky cinquefoil
Caudex branches

elongate.

Stems

openly tufted to loosely spaced, (0.3–)0.6–4 dm;

base 1–3 mm diam., ± densely septate-glandular.

0.6–2.5 dm, base 1–2(–3) mm diam., short hairs sparse to moderately abundant, sometimes absent.
Leaves

glabrate or sparsely to ± densely hairy;

basal (2–)3–16 cm, leaflet pairs (2–)3–4(–5);

terminal leaflet broadly obovate-cuneate to flabellate, 0.2–3(–4) × 0.5–3 cm, teeth single or double, 2–15 per side, apex usually rounded to truncate, sometimes obtuse;

cauline 0–2, reduced, leaflet pairs 2–3.

Basal leaves

3–9(–15) cm, sparsely to densely hairy (hairs to 1.5 mm), usually densely peglike-glandular, not bristly;

leaflet pairs (2–)3(–4);

terminal leaflet broadly obovate-cuneate, 0.8–2(–4) × 0.7–2(–3) cm, teeth single or ± double, 3–8(–12) per side.
Inflorescences

2–40-flowered, not or ± leafy, open, 1/6–3/4(–4/5) of stem, ± wide, branch angles (10–)20–40(–50)°.

Pedicels

3–20 (proximal to 40) mm, not or sparsely to moderately short-hairy, predominantly septate-glandular.

3–15 (proximal to 20) mm, usually not bristly, short hairs sparse to moderately abundant (sometimes absent).
Flowers

opening widely;

epicalyx bractlets linear to elliptic, 2–6 × 1–2 mm;

sepals spreading, 4–7(–9) mm, apex acute to obtuse, apiculate;

petals overlapping or not, spreading, cream-white to pale yellow (red-tinged in var. crumiana), narrowly to broadly obovate, 4–12 × 3–11 mm, longer than sepals;

filaments 1–4 mm, anthers 0.7–1.2 mm;

styles thickened, 1–1.5 mm.

(2–)3–12(–20);

hypanthia and sepals not bristly or bristles less than 1 mm;

epicalyx bractlets linear-lanceolate to broadly elliptic, 2–5 × 1–1.5 mm;

sepals 4–6(–7) mm;

petals ± overlapping, not red-tinged, narrowly to broadly obovate, 4–8(–9) × 3–6(–8) mm;

filaments 1–2.5 mm;

styles usually golden brown, rarely reddish.
Achenes

light brown, 1 mm.

Drymocallis pseudorupestris

Drymocallis pseudorupestris var. saxicola
Phenology Flowering May–Aug.
Habitat Cliffs, ledges, outcrops, ridges, talus slopes, lava beds, other rocky habitats
Elevation 1000–3400 m (3300–11200 ft)
Distribution
from FNA
CA; ID; MT; NV; OR; UT; WA; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; ID; MT; NV; OR; UT; WA; WY; AB; BC
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

Drymocallis pseudorupestris occurs from Alberta and Washington to California and Utah, mostly in montane habitats; it is the species most often associated with rocky habitats, including talus slopes, for which its relatively elongate caudex branches are an obvious adaptation. Vestiture is dominated by abundant septate glands on stems and in the inflorescences. Except for var. pseudorupestris, which occurs only in the northeastern part of the species range, plants are relatively short, usually less than 2.5 dm. Three intergrading varieties accommodate the extremes at the northeastern and southern ends of the range.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Variety saxicola accommodates the bulk of specimens previously placed in Potentilla glandulosa var. pseudorupestris, minus the extremes at the northeastern and southern ends of the species range. The circumscription here encompasses significant heterogeneity, which might be resolved into additional taxa upon further analysis [for example, the dwarfed high elevation form of P. glandulosa noted by N. H. Holmgren (1997b)]. Some collections from Alberta and Washington have petals as large as those of var. pseudorupestris but are here included in var. saxicola on the basis of stature. Plants from southwestern Idaho and southeastern Oregon combine features of D. lactea and D. pseudorupestris, some being atypically tall (to 5.5 dm) but with the vestiture and saxicolous preference of var. saxicola. In contrast, populations on Steens Mountain, Oregon, including the type of D. pumila Rydberg, have the typical habit of var. saxicola but the vestiture of D. lactea; their optimal placement is unresolved. In California, var. saxicola occurs in the Cascade Range (Mount Lassen, Mount Shasta) and extends sporadically south through the Sierra Nevada to Tulare County, intergrading with var. crumiana and D. lactea.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Key
1. Stems (1–)2–4 dm, bases (1.5–)2–3 mm diam.; basal leaves (4–)7–16 cm; terminal leaflets (1–)2–3(–4) cm, teeth usually double, (5–)8–15 per side; flowers (5–)10–40; petals 6–12 × 5–11 mm, widely overlapping; filaments 2–4 mm.
var. pseudorupestris
1. Stems (0.3–)0.6–2.5 dm, bases 1–2(–3) mm diam.; basal leaves (2–)3–9(–15) cm; terminal leaflets 0.2–2(–4) cm, teeth single or ± double, 2–8(–12) per side; flowers 2–12(–20); petals 4–8(–9) × 3–6(–8) mm, not or ± overlapping; filaments 1–2.5(–3) mm
→ 2
2. Basal leaves: leaflet pairs (2–)3(–4); hypanthia and sepals not bristly or bristles less than 1 mm; short hairs sparse to moderately abundant on stems and pedicels (sometimes absent); styles usually golden brown, rarely reddish.
var. saxicola
2. Basal leaves: leaflet pairs 3–4(–5); hypanthia and sepals prominently bristly, bristles 1–1.5 mm; short hairs absent or sparse on stems and pedicels; styles usually dark red, rarely golden brown.
var. crumiana
Source FNA vol. 9, p. 289. FNA vol. 9, p. 290.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis > Drymocallis pseudorupestris
Sibling taxa
D. arguta, D. arizonica, D. ashlandica, D. campanulata, D. convallaria, D. cuneifolia, D. deseretica, D. fissa, D. glabrata, D. glandulosa, D. hansenii, D. lactea, D. micropetala, D. rhomboidea
D. pseudorupestris var. crumiana, D. pseudorupestris var. pseudorupestris
Subordinate taxa
D. pseudorupestris var. crumiana, D. pseudorupestris var. pseudorupestris, D. pseudorupestris var. saxicola
Synonyms Potentilla pseudorupestris, D. glandulosa subsp. pseudorupestris, P. glandulosa subsp. pseudorupestris, P. glandulosa var. pseudorupestris, P. rupestris var. americana
Name authority (Rydberg) Rydberg: Monogr. N. Amer. Potentilleae, 194. (1898) Ertter: J. Bot. Res. Inst. Texas 1: 37, figs. 1M–R. (2007)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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