Flora of North America species comparison

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Drymocallis pseudorupestris

Drymocallis glandulosa
Drymocallis pseudorupestris

cliff drymocallis, cliff woodbeauty, false rock loving cinquefoil, Rocky Mountain sticky cinquefoil

Douglas' woodbeauty, Greene's drymocallis, sticky cinquefoil
Caudex branches

elongate.

short.
Stems

openly tufted to loosely spaced, (0.3–)0.6–4 dm;

base 1–3 mm diam., ± densely septate-glandular.

tufted, (1–)1.5–7.5 dm;

base 1–5 mm diam., moderately to densely septate-glandular.
Leaves

glabrate or sparsely to ± densely hairy;

basal (2–)3–16 cm, leaflet pairs (2–)3–4(–5);

terminal leaflet broadly obovate-cuneate to flabellate, 0.2–3(–4) × 0.5–3 cm, teeth single or double, 2–15 per side, apex usually rounded to truncate, sometimes obtuse;

cauline 0–2, reduced, leaflet pairs 2–3.

sparsely to moderately hairy;

basal (3–)6–30 cm, leaflet pairs (1–)2–4;

terminal leaflet broadly obovate to rhombic-elliptic, (1–)1.5–7(–8) × 1–5 cm, teeth usually double (irregularly single in var. viscida), (4–)6–18(–20) per side, apex rounded to obtuse;

cauline 0–3, developed or reduced, leaflet pairs 1–4.
Inflorescences

2–40-flowered, not or ± leafy, open, 1/6–3/4(–4/5) of stem, ± wide, branch angles (10–)20–40(–50)°.

(2–)5–40-flowered, leafy or not, open or of congested clusters terminating branches, (1/10–)1/5–3/4 of stem, usually wide (narrow in var. viscida), branch angles (15–)20–55°.
Pedicels

3–20 (proximal to 40) mm, not or sparsely to moderately short-hairy, predominantly septate-glandular.

1–10 (proximal to 30) mm, not or sparsely short-hairy, predominantly septate-glandular.
Flowers

opening widely;

epicalyx bractlets linear to elliptic, 2–6 × 1–2 mm;

sepals spreading, 4–7(–9) mm, apex acute to obtuse, apiculate;

petals overlapping or not, spreading, cream-white to pale yellow (red-tinged in var. crumiana), narrowly to broadly obovate, 4–12 × 3–11 mm, longer than sepals;

filaments 1–4 mm, anthers 0.7–1.2 mm;

styles thickened, 1–1.5 mm.

opening widely;

epicalyx bractlets linear-lanceolate to elliptic, 1.5–7(–8) × 0.5–2.5(–3) mm;

sepals spreading or reflexed, (3–)4–11 mm, apex usually broadly obtuse with mucronate tip, sometimes acute;

petals not overlapping, spreading or reflexed, cream-white to yellow, narrowly to broadly obovate or ovate to nearly round, 2–6.5 × 1.5–5.5 mm, shorter than or equal to sepals;

filaments 0.5–2 mm, anthers 0.5–1 mm;

styles thickened, 1–1.5 mm.
Achenes

light brown, 1 mm.

reddish brown, 0.9–1.5 mm.

Drymocallis pseudorupestris

Drymocallis glandulosa
Distribution
from FNA
CA; ID; MT; NV; OR; UT; WA; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; ID; MT; NV; OR; UT; WA; BC; nw Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

Drymocallis pseudorupestris occurs from Alberta and Washington to California and Utah, mostly in montane habitats; it is the species most often associated with rocky habitats, including talus slopes, for which its relatively elongate caudex branches are an obvious adaptation. Vestiture is dominated by abundant septate glands on stems and in the inflorescences. Except for var. pseudorupestris, which occurs only in the northeastern part of the species range, plants are relatively short, usually less than 2.5 dm. Three intergrading varieties accommodate the extremes at the northeastern and southern ends of the range.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 4 (4 in the flora).

Even with the removal of most of the subspecies of Potentilla glandulosa treated by Keck, Drymocallis glandulosa encompasses an inordinate range of variation. This is because of the apparently clinal variation between the coastal extreme (var. wrangelliana), with leafy inflorescences and broad cream-white petals, the montane var. reflexa, with less leafy inflorescences and narrow yellow petals, and the transitional interior var. glandulosa. Variety reflexa, in turn, intergrades in the mountains of southern California with var. viscida, which is anomalous in D. glandulosa with its narrow inflorescences and irregularly single-toothed leaflets. Variation outside of California is unclear, with no evident pattern to differences in petal size and color; these have largely been assigned to var. glandulosa.

At its most distinctive, Drymocallis glandulosa is a robust, densely glandular plant with widely branched leafy inflorescences (except var. viscida), relatively short, stout pedicels, broad sepals with rounded, mucronate tips (except var. viscida and sometimes var. reflexa) that are often significantly enlarged in fruit, and non-overlapping petals that are shorter than to scarcely equal to the sepals. The relatively large achenes are blunter and redder than those of most other species of Drymocallis, as discussed by J. Clausen et al. (1940).

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Key
1. Stems (1–)2–4 dm, bases (1.5–)2–3 mm diam.; basal leaves (4–)7–16 cm; terminal leaflets (1–)2–3(–4) cm, teeth usually double, (5–)8–15 per side; flowers (5–)10–40; petals 6–12 × 5–11 mm, widely overlapping; filaments 2–4 mm.
var. pseudorupestris
1. Stems (0.3–)0.6–2.5 dm, bases 1–2(–3) mm diam.; basal leaves (2–)3–9(–15) cm; terminal leaflets 0.2–2(–4) cm, teeth single or ± double, 2–8(–12) per side; flowers 2–12(–20); petals 4–8(–9) × 3–6(–8) mm, not or ± overlapping; filaments 1–2.5(–3) mm
→ 2
2. Basal leaves: leaflet pairs (2–)3(–4); hypanthia and sepals not bristly or bristles less than 1 mm; short hairs sparse to moderately abundant on stems and pedicels (sometimes absent); styles usually golden brown, rarely reddish.
var. saxicola
2. Basal leaves: leaflet pairs 3–4(–5); hypanthia and sepals prominently bristly, bristles 1–1.5 mm; short hairs absent or sparse on stems and pedicels; styles usually dark red, rarely golden brown.
var. crumiana
1. Petals obovate-elliptic to broadly ovate to nearly round, (2–)3.5–5.5 mm wide, cream-white to ± yellow; sepals spreading, apices obtuse to rounded; inflorescences ± to very leafy; pedicels 1–5 (proximal to 30) mm
→ 2
1. Petals narrowly ovate to narrowly obovate or obovate-elliptic, 1.5–3 mm wide, yellow; sepals reflexed, apices acute to obtuse; inflorescences ± leafy or not; pedicels 2–10 (proximal to 20) mm
→ 3
2. Petals (3–)3.5–5 mm, usually ± yellow, sometimes cream-white; inland.
var. glandulosa
2. Petals (4–)4.5–6.5 mm, usually cream-white, sometimes pale yellow; ± coastal.
var. wrangelliana
3. Inflorescences usually wide, branch angles (15–)25–55°; leaflet teeth usually double, rarely single.
var. reflexa
3. Inflorescences narrow, branch angles 15–30(–40)°; leaflet teeth single or double.
var. viscida
Source FNA vol. 9, p. 289. FNA vol. 9, p. 292.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis
Sibling taxa
D. arguta, D. arizonica, D. ashlandica, D. campanulata, D. convallaria, D. cuneifolia, D. deseretica, D. fissa, D. glabrata, D. glandulosa, D. hansenii, D. lactea, D. micropetala, D. rhomboidea
D. arguta, D. arizonica, D. ashlandica, D. campanulata, D. convallaria, D. cuneifolia, D. deseretica, D. fissa, D. glabrata, D. hansenii, D. lactea, D. micropetala, D. pseudorupestris, D. rhomboidea
Subordinate taxa
D. pseudorupestris var. crumiana, D. pseudorupestris var. pseudorupestris, D. pseudorupestris var. saxicola
D. glandulosa var. glandulosa, D. glandulosa var. reflexa, D. glandulosa var. viscida, D. glandulosa var. wrangelliana
Synonyms Potentilla pseudorupestris, D. glandulosa subsp. pseudorupestris, P. glandulosa subsp. pseudorupestris, P. glandulosa var. pseudorupestris, P. rupestris var. americana Potentilla glandulosa
Name authority (Rydberg) Rydberg: Monogr. N. Amer. Potentilleae, 194. (1898) (Lindley) Rydberg: Monogr. N. Amer. Potentilleae, 198. (1898)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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