Drymocallis pseudorupestris |
Drymocallis glandulosa |
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cliff drymocallis, cliff woodbeauty, false rock loving cinquefoil, Rocky Mountain sticky cinquefoil |
Douglas' woodbeauty, Greene's drymocallis, sticky cinquefoil |
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Caudex branches | elongate. |
short. |
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Stems | openly tufted to loosely spaced, (0.3–)0.6–4 dm; base 1–3 mm diam., ± densely septate-glandular. |
tufted, (1–)1.5–7.5 dm; base 1–5 mm diam., moderately to densely septate-glandular. |
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Leaves | glabrate or sparsely to ± densely hairy; basal (2–)3–16 cm, leaflet pairs (2–)3–4(–5); terminal leaflet broadly obovate-cuneate to flabellate, 0.2–3(–4) × 0.5–3 cm, teeth single or double, 2–15 per side, apex usually rounded to truncate, sometimes obtuse; cauline 0–2, reduced, leaflet pairs 2–3. |
sparsely to moderately hairy; basal (3–)6–30 cm, leaflet pairs (1–)2–4; terminal leaflet broadly obovate to rhombic-elliptic, (1–)1.5–7(–8) × 1–5 cm, teeth usually double (irregularly single in var. viscida), (4–)6–18(–20) per side, apex rounded to obtuse; cauline 0–3, developed or reduced, leaflet pairs 1–4. |
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Inflorescences | 2–40-flowered, not or ± leafy, open, 1/6–3/4(–4/5) of stem, ± wide, branch angles (10–)20–40(–50)°. |
(2–)5–40-flowered, leafy or not, open or of congested clusters terminating branches, (1/10–)1/5–3/4 of stem, usually wide (narrow in var. viscida), branch angles (15–)20–55°. |
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Pedicels | 3–20 (proximal to 40) mm, not or sparsely to moderately short-hairy, predominantly septate-glandular. |
1–10 (proximal to 30) mm, not or sparsely short-hairy, predominantly septate-glandular. |
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Flowers | opening widely; epicalyx bractlets linear to elliptic, 2–6 × 1–2 mm; sepals spreading, 4–7(–9) mm, apex acute to obtuse, apiculate; petals overlapping or not, spreading, cream-white to pale yellow (red-tinged in var. crumiana), narrowly to broadly obovate, 4–12 × 3–11 mm, longer than sepals; filaments 1–4 mm, anthers 0.7–1.2 mm; styles thickened, 1–1.5 mm. |
opening widely; epicalyx bractlets linear-lanceolate to elliptic, 1.5–7(–8) × 0.5–2.5(–3) mm; sepals spreading or reflexed, (3–)4–11 mm, apex usually broadly obtuse with mucronate tip, sometimes acute; petals not overlapping, spreading or reflexed, cream-white to yellow, narrowly to broadly obovate or ovate to nearly round, 2–6.5 × 1.5–5.5 mm, shorter than or equal to sepals; filaments 0.5–2 mm, anthers 0.5–1 mm; styles thickened, 1–1.5 mm. |
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Achenes | light brown, 1 mm. |
reddish brown, 0.9–1.5 mm. |
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Drymocallis pseudorupestris |
Drymocallis glandulosa |
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Distribution |
CA; ID; MT; NV; OR; UT; WA; WY; AB; BC
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AZ; CA; ID; MT; NV; OR; UT; WA; BC; nw Mexico
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Discussion | Varieties 3 (3 in the flora). Drymocallis pseudorupestris occurs from Alberta and Washington to California and Utah, mostly in montane habitats; it is the species most often associated with rocky habitats, including talus slopes, for which its relatively elongate caudex branches are an obvious adaptation. Vestiture is dominated by abundant septate glands on stems and in the inflorescences. Except for var. pseudorupestris, which occurs only in the northeastern part of the species range, plants are relatively short, usually less than 2.5 dm. Three intergrading varieties accommodate the extremes at the northeastern and southern ends of the range. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 4 (4 in the flora). Even with the removal of most of the subspecies of Potentilla glandulosa treated by Keck, Drymocallis glandulosa encompasses an inordinate range of variation. This is because of the apparently clinal variation between the coastal extreme (var. wrangelliana), with leafy inflorescences and broad cream-white petals, the montane var. reflexa, with less leafy inflorescences and narrow yellow petals, and the transitional interior var. glandulosa. Variety reflexa, in turn, intergrades in the mountains of southern California with var. viscida, which is anomalous in D. glandulosa with its narrow inflorescences and irregularly single-toothed leaflets. Variation outside of California is unclear, with no evident pattern to differences in petal size and color; these have largely been assigned to var. glandulosa. At its most distinctive, Drymocallis glandulosa is a robust, densely glandular plant with widely branched leafy inflorescences (except var. viscida), relatively short, stout pedicels, broad sepals with rounded, mucronate tips (except var. viscida and sometimes var. reflexa) that are often significantly enlarged in fruit, and non-overlapping petals that are shorter than to scarcely equal to the sepals. The relatively large achenes are blunter and redder than those of most other species of Drymocallis, as discussed by J. Clausen et al. (1940). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 289. | FNA vol. 9, p. 292. | ||||||||||||||||||||
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis | ||||||||||||||||||||
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Synonyms | Potentilla pseudorupestris, D. glandulosa subsp. pseudorupestris, P. glandulosa subsp. pseudorupestris, P. glandulosa var. pseudorupestris, P. rupestris var. americana | Potentilla glandulosa | ||||||||||||||||||||
Name authority | (Rydberg) Rydberg: Monogr. N. Amer. Potentilleae, 194. (1898) | (Lindley) Rydberg: Monogr. N. Amer. Potentilleae, 198. (1898) | ||||||||||||||||||||
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