Drymocallis pseudorupestris |
Drymocallis ashlandica |
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cliff drymocallis, cliff woodbeauty, false rock loving cinquefoil, Rocky Mountain sticky cinquefoil |
Ashland cinquefoil, Mazama drymocallis or wood beauty |
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Caudex branches | elongate. |
short. |
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Stems | openly tufted to loosely spaced, (0.3–)0.6–4 dm; base 1–3 mm diam., ± densely septate-glandular. |
tufted, (1–)2–5 dm; base 2–3 mm diam., ± densely septate-glandular. |
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Leaves | glabrate or sparsely to ± densely hairy; basal (2–)3–16 cm, leaflet pairs (2–)3–4(–5); terminal leaflet broadly obovate-cuneate to flabellate, 0.2–3(–4) × 0.5–3 cm, teeth single or double, 2–15 per side, apex usually rounded to truncate, sometimes obtuse; cauline 0–2, reduced, leaflet pairs 2–3. |
moderately to ± densely hairy; basal 6–16 cm, leaflet pairs 2–3(–4); terminal leaflet broadly obovate, 1.5–4 × 1–3 cm, teeth ± double, 7–12 per side, apex rounded; cauline 0–2, moderately developed, leaflet pairs 2–3. |
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Inflorescences | 2–40-flowered, not or ± leafy, open, 1/6–3/4(–4/5) of stem, ± wide, branch angles (10–)20–40(–50)°. |
5–15(–20)-flowered, not leafy, congested to deeply branched, 1/10–2/3 or less of stem, usually narrow, rarely ± wide, branch angles (5–)10–20(–30)°. |
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Pedicels | 3–20 (proximal to 40) mm, not or sparsely to moderately short-hairy, predominantly septate-glandular. |
2–10 (proximal to 30) mm, sparsely to moderately short-hairy, predominantly septate-glandular, often obscuring other vestiture. |
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Flowers | opening widely; epicalyx bractlets linear to elliptic, 2–6 × 1–2 mm; sepals spreading, 4–7(–9) mm, apex acute to obtuse, apiculate; petals overlapping or not, spreading, cream-white to pale yellow (red-tinged in var. crumiana), narrowly to broadly obovate, 4–12 × 3–11 mm, longer than sepals; filaments 1–4 mm, anthers 0.7–1.2 mm; styles thickened, 1–1.5 mm. |
opening widely; epicalyx bractlets linear to broadly elliptic, 3–5 × 1–2 mm; sepals spreading, 5–7(–9) mm, apex acute; petals overlapping, spreading, light yellow, broadly obovate, 5–9 × 4–7 mm, equal to or longer than sepals; filaments 1.5–5 mm, anthers 0.7–1.2 mm; styles ± thickened, 1 mm. |
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Achenes | light brown, 1 mm. |
light brown, 1 mm. |
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Drymocallis pseudorupestris |
Drymocallis ashlandica |
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Phenology | Flowering Jun–Aug. | |||||||||
Habitat | Meadows, roadsides, sandy shorelines | |||||||||
Elevation | 500–2000 m (1600–6600 ft) | |||||||||
Distribution |
CA; ID; MT; NV; OR; UT; WA; WY; AB; BC
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OR
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Discussion | Varieties 3 (3 in the flora). Drymocallis pseudorupestris occurs from Alberta and Washington to California and Utah, mostly in montane habitats; it is the species most often associated with rocky habitats, including talus slopes, for which its relatively elongate caudex branches are an obvious adaptation. Vestiture is dominated by abundant septate glands on stems and in the inflorescences. Except for var. pseudorupestris, which occurs only in the northeastern part of the species range, plants are relatively short, usually less than 2.5 dm. Three intergrading varieties accommodate the extremes at the northeastern and southern ends of the range. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The name Potentilla glandulosa subsp. ashlandica was misapplied to Drymocallis lactea var. austiniae (B. Ertter 1993); as circumscribed here, D. ashlandica is a poorly known species restricted to the Cascade and Siskiyou ranges of southwestern Oregon. It very likely also occurs in adjacent California; no collections from there are known. Drymocallis ashlandica differs from most sympatric populations of D. lactea var. austiniae and resembles D. pseudorupestris in its densely glandular inflorescences, which tend to be more densely congested and/or narrower than those of the other two species. Drymocallis ashlandica was described by Greene as Potentilla ashlandica, and by Howell as P. ciliata (a later homonym), both based on a collection by Howell from the Siskiyou Mountains near Ashland, Oregon. The circumscription here, which matches that of D. D. Keck (in J. Clausen et al. 1940) and M. E. Peck (1961), encompasses populations from near Oregon Caves and Ashland in the Siskiyou Mountains to near McKenzie Bridge, Lane County, and Sparks Lake, Deschutes County. It is currently known from relatively few collections, mostly from the Crater Lake area, and may prove to be of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 289. | FNA vol. 9, p. 288. | ||||||||
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis | ||||||||
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Synonyms | Potentilla pseudorupestris, D. glandulosa subsp. pseudorupestris, P. glandulosa subsp. pseudorupestris, P. glandulosa var. pseudorupestris, P. rupestris var. americana | Potentilla ashlandica, D. glandulosa subsp. ashlandica, P. glandulosa subsp. ashlandica | ||||||||
Name authority | (Rydberg) Rydberg: Monogr. N. Amer. Potentilleae, 194. (1898) | (Greene) Rydberg: Monogr. N. Amer. Potentilleae, 200. (1898) | ||||||||
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