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Drymocallis pseudorupestris

cliff drymocallis, cliff woodbeauty, false rock loving cinquefoil, Rocky Mountain sticky cinquefoil

Arizona drymocallis or wood beauty

Caudex branches

elongate.

short.

Stems

openly tufted to loosely spaced, (0.3–)0.6–4 dm;

base 1–3 mm diam., ± densely septate-glandular.

± tufted, (1.5–)2.5–6 dm;

base 1.5–2.5 mm diam., sparsely to densely septate-glandular.

Leaves

glabrate or sparsely to ± densely hairy;

basal (2–)3–16 cm, leaflet pairs (2–)3–4(–5);

terminal leaflet broadly obovate-cuneate to flabellate, 0.2–3(–4) × 0.5–3 cm, teeth single or double, 2–15 per side, apex usually rounded to truncate, sometimes obtuse;

cauline 0–2, reduced, leaflet pairs 2–3.

sparsely to moderately hairy;

basal 5–22 cm, leaflet pairs (2–)3–4(–5);

terminal leaflet broadly obovate-elliptic, 1–5 × 1–2.5(–4) cm, teeth double, 6–18 per side, apex rounded to acute;

cauline 1–4, well developed, leaflet pairs 2–3.

Inflorescences

2–40-flowered, not or ± leafy, open, 1/6–3/4(–4/5) of stem, ± wide, branch angles (10–)20–40(–50)°.

5–30-flowered, not notably leafy, compact, 1/10–1/4(–1/2) of stem, narrow, branch angles 5–25°.

Pedicels

3–20 (proximal to 40) mm, not or sparsely to moderately short-hairy, predominantly septate-glandular.

1–10 (proximal to 20) mm, sparsely to ± densely short-hairy, predominantly septate-glandular.

Flowers

opening widely;

epicalyx bractlets linear to elliptic, 2–6 × 1–2 mm;

sepals spreading, 4–7(–9) mm, apex acute to obtuse, apiculate;

petals overlapping or not, spreading, cream-white to pale yellow (red-tinged in var. crumiana), narrowly to broadly obovate, 4–12 × 3–11 mm, longer than sepals;

filaments 1–4 mm, anthers 0.7–1.2 mm;

styles thickened, 1–1.5 mm.

opening widely;

epicalyx bractlets linear-lanceolate to narrowly elliptic, 2–4(–7) × 1–2 mm;

sepals spreading, 4–6(–11) mm, apex ± acute;

petals not or scarcely overlapping, spreading, cream-white, ± obovate, 3–6 × 2.5–5 mm, shorter than or equal to sepals;

filaments 1–3 mm, anthers 0.7–1 mm;

styles thickened, 1 mm.

Achenes

light brown, 1 mm.

light brown, 1 mm.

2n

= 14.

Drymocallis pseudorupestris

Drymocallis arizonica

Phenology Flowering May–Jul.
Habitat Streamsides, rocky sites, open forest floors, pine and aspen forests
Elevation 1800–3200 m (5900–10500 ft)
Distribution
from FNA
CA; ID; MT; NV; OR; UT; WA; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; UT
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

Drymocallis pseudorupestris occurs from Alberta and Washington to California and Utah, mostly in montane habitats; it is the species most often associated with rocky habitats, including talus slopes, for which its relatively elongate caudex branches are an obvious adaptation. Vestiture is dominated by abundant septate glands on stems and in the inflorescences. Except for var. pseudorupestris, which occurs only in the northeastern part of the species range, plants are relatively short, usually less than 2.5 dm. Three intergrading varieties accommodate the extremes at the northeastern and southern ends of the range.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Drymocallis arizonica encompasses populations in Arizona north of the Mogollon Rim and in Utah as far north as Garfield and Sevier counties, including the Henry Mountains. It is most distinctive in northern Arizona, where plants commonly have basal leaves with four pairs of lateral leaflets and compact inflorescences. The species intergrades with D. convallaria but is in general shorter and more likely multistemmed. It also shares similarities with D. pseudorupestris but has more developed cauline leaves and a more compact inflorescence. The intergrade zone between all three species and D. deseretica in southwestern Utah is particularly problematic and perhaps indicative of yet additional taxa deserving recognition.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems (1–)2–4 dm, bases (1.5–)2–3 mm diam.; basal leaves (4–)7–16 cm; terminal leaflets (1–)2–3(–4) cm, teeth usually double, (5–)8–15 per side; flowers (5–)10–40; petals 6–12 × 5–11 mm, widely overlapping; filaments 2–4 mm.
var. pseudorupestris
1. Stems (0.3–)0.6–2.5 dm, bases 1–2(–3) mm diam.; basal leaves (2–)3–9(–15) cm; terminal leaflets 0.2–2(–4) cm, teeth single or ± double, 2–8(–12) per side; flowers 2–12(–20); petals 4–8(–9) × 3–6(–8) mm, not or ± overlapping; filaments 1–2.5(–3) mm
→ 2
2. Basal leaves: leaflet pairs (2–)3(–4); hypanthia and sepals not bristly or bristles less than 1 mm; short hairs sparse to moderately abundant on stems and pedicels (sometimes absent); styles usually golden brown, rarely reddish.
var. saxicola
2. Basal leaves: leaflet pairs 3–4(–5); hypanthia and sepals prominently bristly, bristles 1–1.5 mm; short hairs absent or sparse on stems and pedicels; styles usually dark red, rarely golden brown.
var. crumiana
Source FNA vol. 9, p. 289. FNA vol. 9, p. 284.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis
Sibling taxa
D. arguta, D. arizonica, D. ashlandica, D. campanulata, D. convallaria, D. cuneifolia, D. deseretica, D. fissa, D. glabrata, D. glandulosa, D. hansenii, D. lactea, D. micropetala, D. rhomboidea
D. arguta, D. ashlandica, D. campanulata, D. convallaria, D. cuneifolia, D. deseretica, D. fissa, D. glabrata, D. glandulosa, D. hansenii, D. lactea, D. micropetala, D. pseudorupestris, D. rhomboidea
Subordinate taxa
D. pseudorupestris var. crumiana, D. pseudorupestris var. pseudorupestris, D. pseudorupestris var. saxicola
Synonyms Potentilla pseudorupestris, D. glandulosa subsp. pseudorupestris, P. glandulosa subsp. pseudorupestris, P. glandulosa var. pseudorupestris, P. rupestris var. americana D. glandulosa subsp. arizonica, Potentilla glandulosa subsp. arizonica, P. macdougalii
Name authority (Rydberg) Rydberg: Monogr. N. Amer. Potentilleae, 194. (1898) Rydberg: in N. L. Britton et al., N. Amer. Fl. 22: 373. (1908)
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