Drymocallis lactea |
Rosaceae tribe potentilleae |
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Nevada cinquefoil, Sierran woodbeauty |
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Habit | Herbs, perennial, rarely annual or biennial, shrubs, or subshrubs; unarmed. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Caudex branches | short. |
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Stems | tufted, (0.3–)1.5–6(–6.5) dm; base 1–2.5 mm diam., not or sparsely, sometimes moderately, septate-glandular. |
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Leaves | usually sparsely to moderately hairy, sometimes glabrate; basal (2–)5–20(–30) cm, leaflet pairs (2–)3–4(–5); terminal leaflet broadly obovate to cuneate, 1–4 × 0.7–3.5 cm, teeth usually single, 4–10(–14) per side, apex usually rounded to obtuse, rarely acute; cauline 1–2, reduced, leaflet pairs 2–3. |
alternate, rarely opposite, pinnately (palmately) compound (simple in Alchemilla, Aphanes, and Chamaerhodos); stipules persistent (absent in Chamaerhodos), adnate to petiole; venation pinnate or palmate. |
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Inflorescences | (2–)5–30(–50)-flowered, not leafy, open, (1/6–)1/5–1/2(–4/5) of stem, narrow to wide, branch angles 10–40(–50)°. |
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Pedicels | 2–10 (proximal to 30) mm, predominantly short-hairy, sometimes velutinous, not or sparsely, sometimes moderately, septate-glandular. |
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Flowers | opening widely; epicalyx bractlets linear-lanceolate to narrowly elliptic, 2–5 × 0.5–1.5 mm; sepals spreading, 3–8(–9) mm, apex usually acute, sometimes obtuse and apiculate; petals ± overlapping, spreading, cream-white to bright yellow, broadly obovate, (3–)4–8(–10) × 3–8 mm, usually longer than, rarely equal to, sepals; filaments 1–3(–3.5) mm, anthers 0.6–1 mm; styles thickened, 1 mm. |
perianth and androecium perigynous; epicalyx bractlets present, sometimes absent; hypanthium usually patelliform, cupulate, or campanulate, sometimes turbinate, saucer-shaped, flat-bottomed, or subglobose to ellipsoid or ovoid; torus flat to conic or turbinate, enlarged (absent or reduced in Alchemilla, Aphanes, and Chamaerhodos); carpels 1–260, styles basal or lateral to subterminal, distinct; ovules 1(or 2), basal. |
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Fruits | aggregated achenes (achenes in Alchemilla and Aphanes); torus sometimes fleshy; styles deciduous or persistent, not elongate. |
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Achenes | light brown, 1 mm. |
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Drymocallis lactea |
Rosaceae tribe potentilleae |
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Distribution |
CA; NV; OR
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North America; Mexico; Central America; South America; West Indies; Bermuda; Eurasia; Africa; Atlantic Islands; Pacific Islands; Australia |
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Discussion | Varieties 2 (2 in the flora). Drymocallis lactea has had a well-established identity as Potentilla glandulosa var. (or subsp.) nevadensis; however, the epithet lactea has priority at species rank and avoids possible confusion with P. nevadensis Boissier. D. D. Keck (in J. Clausen et al. 1940) based his concept of P. glandulosa subsp. nevadensis on white-petaled plants with narrow inflorescences, the common form in the central Sierra Nevada and south. His map and annotations, however, include populations with yellow petals and widely branched inflorescences from northern California and adjacent Oregon. The two extremes merge in the northern Sierra Nevada, and populations in the North Coast Ranges of California also are problematic; the extremes are accordingly treated here as varieties of a single species. The inclusion by Keck of Washington in the species range might have been based on specimens here assigned to P. pseudorupestris. Drymocallis lactea can be distinguished from sympatric species by its relatively large, overlapping petals, non-leafy inflorescences, and singly toothed leaflets. Pedicels and distal stems are usually densely short-hairy with relatively few or no glandular hairs (in contrast to D. pseudorupestris), and septate glands are usually absent from stem bases (in contrast to D. hansenii). The distinction between D. lactea and D. pseudorupestris breaks down at the margin of the species range in Nevada and Oregon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 14–22, species ca. 860 (14 genera, 189 species, including 1 hybrid, in the flora area). The base chromosome number for Potentilleae is mostly x = 7 (8 in Alchemilla and Aphanes; 14 in Comarum). Variation in the number of genera recognized in Potentilleae is due to differences in generic delimitation between D. Potter et al. (2007) and the authors of Potentilla and segregates here (see 9. Ivesia and 8. Potentilla for discussion). In the former, Duchesnea, Horkelia, Horkeliella, and Ivesia are included within Potentilla. Likewise, Aphanes is included within Alchemilla by Potter et al. while it is kept distinct here. Potentilla and its segregates and Fragaria are host to Phragmidium rusts, but not the other genera of the tribe. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 287. | FNA vol. 9, p. 119. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis | Rosaceae > subfam. Rosoideae | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | Potentilla glandulosa var. lactea | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | (Greene) Rydberg: in N. L. Britton et al., N. Amer. Fl. 22: 369. (1908) | Sweet: Brit. Fl. Gard. 2: sub plate 124. (1825) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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