Drymocallis lactea
Drymocallis lactea var. austiniae
Nevada cinquefoil, Sierran woodbeauty
Austin's drymocallis, Austin's drymocallis or wood beauty, Austin's woodbeauty
short.
tufted, (0.3–)1.5–6(–6.5) dm;
base 1–2.5 mm diam., not or sparsely, sometimes moderately, septate-glandular.
(0.4–)1.5–4.5(–6.5) dm;
base sparsely to densely short-hairy, sometimes moderately septate-glandular.
usually sparsely to moderately hairy, sometimes glabrate;
basal (2–)5–20(–30) cm, leaflet pairs (2–)3–4(–5);
terminal leaflet broadly obovate to cuneate, 1–4 × 0.7–3.5 cm, teeth usually single, 4–10(–14) per side, apex usually rounded to obtuse, rarely acute;
cauline 1–2, reduced, leaflet pairs 2–3.
(2–)5–30(–50)-flowered, not leafy, open, (1/6–)1/5–1/2(–4/5) of stem, narrow to wide, branch angles 10–40(–50)°.
(2–)5–30(–50)-flowered, (1/6–)1/4–1/2(–4/5) of stem, wide, branch angles (10–)20–40(–50)°.
2–10 (proximal to 30) mm, predominantly short-hairy, sometimes velutinous, not or sparsely, sometimes moderately, septate-glandular.
2–20(–30) mm, sparsely to ± densely short-hairy, usually eglandular, sometimes sparsely to moderately septate-glandular.
opening widely;
epicalyx bractlets linear-lanceolate to narrowly elliptic, 2–5 × 0.5–1.5 mm;
sepals spreading, 3–8(–9) mm, apex usually acute, sometimes obtuse and apiculate;
petals ± overlapping, spreading, cream-white to bright yellow, broadly obovate, (3–)4–8(–10) × 3–8 mm, usually longer than, rarely equal to, sepals;
filaments 1–3(–3.5) mm, anthers 0.6–1 mm;
styles thickened, 1 mm.
light brown, 1 mm.
(cream-white) pale to bright yellow.
Drymocallis lactea
Drymocallis lactea var. austiniae
Varieties 2 (2 in the flora).
Drymocallis lactea has had a well-established identity as Potentilla glandulosa var. (or subsp.) nevadensis; however, the epithet lactea has priority at species rank and avoids possible confusion with P. nevadensis Boissier. D. D. Keck (in J. Clausen et al. 1940) based his concept of P. glandulosa subsp. nevadensis on white-petaled plants with narrow inflorescences, the common form in the central Sierra Nevada and south. His map and annotations, however, include populations with yellow petals and widely branched inflorescences from northern California and adjacent Oregon. The two extremes merge in the northern Sierra Nevada, and populations in the North Coast Ranges of California also are problematic; the extremes are accordingly treated here as varieties of a single species. The inclusion by Keck of Washington in the species range might have been based on specimens here assigned to P. pseudorupestris.
Drymocallis lactea can be distinguished from sympatric species by its relatively large, overlapping petals, non-leafy inflorescences, and singly toothed leaflets. Pedicels and distal stems are usually densely short-hairy with relatively few or no glandular hairs (in contrast to D. pseudorupestris), and septate glands are usually absent from stem bases (in contrast to D. hansenii). The distinction between D. lactea and D. pseudorupestris breaks down at the margin of the species range in Nevada and Oregon.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Variety austiniae was erroneously treated as Potentilla glandulosa subsp. ashlandica (Greene) D. D. Keck (B. Ertter 1993); it now is understood to be a separate entity. As here defined, it is the common large-petaled, open-flowered Drymocallis of northern California and adjacent Oregon and Nevada, differing from var. lactea in its more widely branched inflorescences and usually pale to bright yellow petals. The boundary between the two varieties is unclear, in part because petal color is uncertain on herbarium specimens. Some collections from Steens Mountain, Oregon, are transitional to D. pseudorupestris var. saxicola. Occasional populations in northern California that combine the morphology of var. austiniae with more glandular vestiture on pedicels and stems might indicate introgression with D. ashlandica or D. pseudorupestris.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Inflorescences narrow, branch angles 10–20°; petals cream-white to pale yellowish; pedicels and stems not or sparsely septate-glandular. | var. lactea |
1. Inflorescences wide, branch angles mostly 20–40°; petals usually pale to bright yellow; pedicels and stems sometimes moderately septate-glandular. | var. austiniae |
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