Drymocallis lactea |
Drymocallis campanulata |
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Nevada cinquefoil, Sierran woodbeauty |
John Day drymocallis, John Day drymocallis or wood beauty |
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Caudex branches | short. |
elongate. |
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Stems | tufted, (0.3–)1.5–6(–6.5) dm; base 1–2.5 mm diam., not or sparsely, sometimes moderately, septate-glandular. |
openly tufted to loosely spaced, 1.5–4.5 dm; base 1.2–3.5 mm diam., ± densely septate-glandular. |
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Leaves | usually sparsely to moderately hairy, sometimes glabrate; basal (2–)5–20(–30) cm, leaflet pairs (2–)3–4(–5); terminal leaflet broadly obovate to cuneate, 1–4 × 0.7–3.5 cm, teeth usually single, 4–10(–14) per side, apex usually rounded to obtuse, rarely acute; cauline 1–2, reduced, leaflet pairs 2–3. |
sparsely to ± densely hairy; basal 6–20 cm, leaflet pairs (2–)3–4(–5); terminal leaflet broadly obovate to nearly round, 1–4 × 1–3.5 cm, teeth usually ± double, 5–10 per side, apex rounded; cauline 0–2, well developed, leaflet pairs 2–3. |
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Inflorescences | (2–)5–30(–50)-flowered, not leafy, open, (1/6–)1/5–1/2(–4/5) of stem, narrow to wide, branch angles 10–40(–50)°. |
5–50-flowered, leafy, open, (1/5–)1/2(–4/5) of stem, wide, branch angles 25–50°. |
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Pedicels | 2–10 (proximal to 30) mm, predominantly short-hairy, sometimes velutinous, not or sparsely, sometimes moderately, septate-glandular. |
2–5 (proximal to 10) mm, sparsely to moderately short-hairy, predominantly septate-glandular. |
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Flowers | opening widely; epicalyx bractlets linear-lanceolate to narrowly elliptic, 2–5 × 0.5–1.5 mm; sepals spreading, 3–8(–9) mm, apex usually acute, sometimes obtuse and apiculate; petals ± overlapping, spreading, cream-white to bright yellow, broadly obovate, (3–)4–8(–10) × 3–8 mm, usually longer than, rarely equal to, sepals; filaments 1–3(–3.5) mm, anthers 0.6–1 mm; styles thickened, 1 mm. |
opening narrowly; epicalyx bractlets usually lanceolate, sometimes linear, 3–4 × 1–1.5 mm; sepals ± erect, 5–8 mm, apex acute to obtuse; petals overlapping, ± erect, light yellow, broadly obovate, 5–11 × (3–)5–8 mm, equal to or exceeding sepals; filaments 1.5–3 mm, anthers 1 mm; styles slender, (1–)1.5–2.5 mm. |
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Achenes | light brown, 1 mm. |
light brown, 1 mm. |
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Drymocallis lactea |
Drymocallis campanulata |
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Phenology | Flowering May–Jul. | |||||
Habitat | Basaltic cliffs and talus above streams | |||||
Elevation | 700–1400 m (2300–4600 ft) | |||||
Distribution |
CA; NV; OR
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OR
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Discussion | Varieties 2 (2 in the flora). Drymocallis lactea has had a well-established identity as Potentilla glandulosa var. (or subsp.) nevadensis; however, the epithet lactea has priority at species rank and avoids possible confusion with P. nevadensis Boissier. D. D. Keck (in J. Clausen et al. 1940) based his concept of P. glandulosa subsp. nevadensis on white-petaled plants with narrow inflorescences, the common form in the central Sierra Nevada and south. His map and annotations, however, include populations with yellow petals and widely branched inflorescences from northern California and adjacent Oregon. The two extremes merge in the northern Sierra Nevada, and populations in the North Coast Ranges of California also are problematic; the extremes are accordingly treated here as varieties of a single species. The inclusion by Keck of Washington in the species range might have been based on specimens here assigned to P. pseudorupestris. Drymocallis lactea can be distinguished from sympatric species by its relatively large, overlapping petals, non-leafy inflorescences, and singly toothed leaflets. Pedicels and distal stems are usually densely short-hairy with relatively few or no glandular hairs (in contrast to D. pseudorupestris), and septate glands are usually absent from stem bases (in contrast to D. hansenii). The distinction between D. lactea and D. pseudorupestris breaks down at the margin of the species range in Nevada and Oregon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Drymocallis campanulata is one of the more distinct species in the genus, with its campanulate flowers, relatively large butter yellow petals, and densely glandular pedicels. It occurs primarily in the canyons of the John Day River in Grant and Wheeler counties. The invalidly published Potentilla campanulata D. D. Keck provides a full description and illustration for the subsequent basionym. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 287. | FNA vol. 9, p. 294. | ||||
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Potentilla glandulosa var. lactea | Potentilla glandulosa var. campanulata | ||||
Name authority | (Greene) Rydberg: in N. L. Britton et al., N. Amer. Fl. 22: 369. (1908) | (C. L. Hitchcock) Ertter: J. Bot. Res. Inst. Texas 1: 43. (2007) | ||||
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