Drymocallis lactea
Drymocallis ashlandica
Nevada cinquefoil, Sierran woodbeauty
Ashland cinquefoil, Mazama drymocallis or wood beauty
short.
short.
tufted, (0.3–)1.5–6(–6.5) dm;
base 1–2.5 mm diam., not or sparsely, sometimes moderately, septate-glandular.
tufted, (1–)2–5 dm;
base 2–3 mm diam., ± densely septate-glandular.
usually sparsely to moderately hairy, sometimes glabrate;
basal (2–)5–20(–30) cm, leaflet pairs (2–)3–4(–5);
terminal leaflet broadly obovate to cuneate, 1–4 × 0.7–3.5 cm, teeth usually single, 4–10(–14) per side, apex usually rounded to obtuse, rarely acute;
cauline 1–2, reduced, leaflet pairs 2–3.
moderately to ± densely hairy;
basal 6–16 cm, leaflet pairs 2–3(–4);
terminal leaflet broadly obovate, 1.5–4 × 1–3 cm, teeth ± double, 7–12 per side, apex rounded;
cauline 0–2, moderately developed, leaflet pairs 2–3.
(2–)5–30(–50)-flowered, not leafy, open, (1/6–)1/5–1/2(–4/5) of stem, narrow to wide, branch angles 10–40(–50)°.
5–15(–20)-flowered, not leafy, congested to deeply branched, 1/10–2/3 or less of stem, usually narrow, rarely ± wide, branch angles (5–)10–20(–30)°.
2–10 (proximal to 30) mm, predominantly short-hairy, sometimes velutinous, not or sparsely, sometimes moderately, septate-glandular.
2–10 (proximal to 30) mm, sparsely to moderately short-hairy, predominantly septate-glandular, often obscuring other vestiture.
opening widely;
epicalyx bractlets linear-lanceolate to narrowly elliptic, 2–5 × 0.5–1.5 mm;
sepals spreading, 3–8(–9) mm, apex usually acute, sometimes obtuse and apiculate;
petals ± overlapping, spreading, cream-white to bright yellow, broadly obovate, (3–)4–8(–10) × 3–8 mm, usually longer than, rarely equal to, sepals;
filaments 1–3(–3.5) mm, anthers 0.6–1 mm;
styles thickened, 1 mm.
opening widely;
epicalyx bractlets linear to broadly elliptic, 3–5 × 1–2 mm;
sepals spreading, 5–7(–9) mm, apex acute;
petals overlapping, spreading, light yellow, broadly obovate, 5–9 × 4–7 mm, equal to or longer than sepals;
filaments 1.5–5 mm, anthers 0.7–1.2 mm;
styles ± thickened, 1 mm.
light brown, 1 mm.
light brown, 1 mm.
Drymocallis lactea
Drymocallis ashlandica
Varieties 2 (2 in the flora).
Drymocallis lactea has had a well-established identity as Potentilla glandulosa var. (or subsp.) nevadensis; however, the epithet lactea has priority at species rank and avoids possible confusion with P. nevadensis Boissier. D. D. Keck (in J. Clausen et al. 1940) based his concept of P. glandulosa subsp. nevadensis on white-petaled plants with narrow inflorescences, the common form in the central Sierra Nevada and south. His map and annotations, however, include populations with yellow petals and widely branched inflorescences from northern California and adjacent Oregon. The two extremes merge in the northern Sierra Nevada, and populations in the North Coast Ranges of California also are problematic; the extremes are accordingly treated here as varieties of a single species. The inclusion by Keck of Washington in the species range might have been based on specimens here assigned to P. pseudorupestris.
Drymocallis lactea can be distinguished from sympatric species by its relatively large, overlapping petals, non-leafy inflorescences, and singly toothed leaflets. Pedicels and distal stems are usually densely short-hairy with relatively few or no glandular hairs (in contrast to D. pseudorupestris), and septate glands are usually absent from stem bases (in contrast to D. hansenii). The distinction between D. lactea and D. pseudorupestris breaks down at the margin of the species range in Nevada and Oregon.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
The name Potentilla glandulosa subsp. ashlandica was misapplied to Drymocallis lactea var. austiniae (B. Ertter 1993); as circumscribed here, D. ashlandica is a poorly known species restricted to the Cascade and Siskiyou ranges of southwestern Oregon. It very likely also occurs in adjacent California; no collections from there are known. Drymocallis ashlandica differs from most sympatric populations of D. lactea var. austiniae and resembles D. pseudorupestris in its densely glandular inflorescences, which tend to be more densely congested and/or narrower than those of the other two species.
Drymocallis ashlandica was described by Greene as Potentilla ashlandica, and by Howell as P. ciliata (a later homonym), both based on a collection by Howell from the Siskiyou Mountains near Ashland, Oregon. The circumscription here, which matches that of D. D. Keck (in J. Clausen et al. 1940) and M. E. Peck (1961), encompasses populations from near Oregon Caves and Ashland in the Siskiyou Mountains to near McKenzie Bridge, Lane County, and Sparks Lake, Deschutes County. It is currently known from relatively few collections, mostly from the Crater Lake area, and may prove to be of conservation concern.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Inflorescences narrow, branch angles 10–20°; petals cream-white to pale yellowish; pedicels and stems not or sparsely septate-glandular. | var. lactea |
1. Inflorescences wide, branch angles mostly 20–40°; petals usually pale to bright yellow; pedicels and stems sometimes moderately septate-glandular. | var. austiniae |
CA
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