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Nevada cinquefoil, Sierran woodbeauty

Nevada cinquefoil, Sierran woodbeauty

Caudex branches

short.

Stems

(0.3–)1–6 dm;

base glabrate or sparsely to densely short-hairy, not or sparsely septate-glandular.

tufted, (0.3–)1.5–6(–6.5) dm;

base 1–2.5 mm diam., not or sparsely, sometimes moderately, septate-glandular.

Leaves

usually sparsely to moderately hairy, sometimes glabrate;

basal (2–)5–20(–30) cm, leaflet pairs (2–)3–4(–5);

terminal leaflet broadly obovate to cuneate, 1–4 × 0.7–3.5 cm, teeth usually single, 4–10(–14) per side, apex usually rounded to obtuse, rarely acute;

cauline 1–2, reduced, leaflet pairs 2–3.

Inflorescences

3–20-flowered, 1/5–2/5(–3/4) of stem, narrow, branch angles 10–20°.

(2–)5–30(–50)-flowered, not leafy, open, (1/6–)1/5–1/2(–4/5) of stem, narrow to wide, branch angles 10–40(–50)°.

Pedicels

5–10 mm, moderately to densely short-hairy, not or sparsely septate-glandular.

2–10 (proximal to 30) mm, predominantly short-hairy, sometimes velutinous, not or sparsely, sometimes moderately, septate-glandular.

Flowers

opening widely;

epicalyx bractlets linear-lanceolate to narrowly elliptic, 2–5 × 0.5–1.5 mm;

sepals spreading, 3–8(–9) mm, apex usually acute, sometimes obtuse and apiculate;

petals ± overlapping, spreading, cream-white to bright yellow, broadly obovate, (3–)4–8(–10) × 3–8 mm, usually longer than, rarely equal to, sepals;

filaments 1–3(–3.5) mm, anthers 0.6–1 mm;

styles thickened, 1 mm.

Achenes

light brown, 1 mm.

Petals

cream-white to pale yellowish.

2n

= 14.

Drymocallis lactea var. lactea

Drymocallis lactea

Phenology Flowering May–Aug.
Habitat Rocky, seasonally moist places
Elevation 1600–3700 m (5200–12100 ft)
Distribution
from FNA
CA; NV; OR
from FNA
CA; NV; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Variety lactea is the most common and conspicuous Drymocallis at higher elevations in the central and southern Sierra Nevada, where it is recognized by its large cream-white to yellowish petals and narrow inflorescences. It also occurs in the San Jacinto Mountains and Transverse Ranges of southern California, and extends into the Intermountain Region from the mountains of southeastern Oregon to central Nevada.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Drymocallis lactea has had a well-established identity as Potentilla glandulosa var. (or subsp.) nevadensis; however, the epithet lactea has priority at species rank and avoids possible confusion with P. nevadensis Boissier. D. D. Keck (in J. Clausen et al. 1940) based his concept of P. glandulosa subsp. nevadensis on white-petaled plants with narrow inflorescences, the common form in the central Sierra Nevada and south. His map and annotations, however, include populations with yellow petals and widely branched inflorescences from northern California and adjacent Oregon. The two extremes merge in the northern Sierra Nevada, and populations in the North Coast Ranges of California also are problematic; the extremes are accordingly treated here as varieties of a single species. The inclusion by Keck of Washington in the species range might have been based on specimens here assigned to P. pseudorupestris.

Drymocallis lactea can be distinguished from sympatric species by its relatively large, overlapping petals, non-leafy inflorescences, and singly toothed leaflets. Pedicels and distal stems are usually densely short-hairy with relatively few or no glandular hairs (in contrast to D. pseudorupestris), and septate glands are usually absent from stem bases (in contrast to D. hansenii). The distinction between D. lactea and D. pseudorupestris breaks down at the margin of the species range in Nevada and Oregon.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Inflorescences narrow, branch angles 10–20°; petals cream-white to pale yellowish; pedicels and stems not or sparsely septate-glandular.
var. lactea
1. Inflorescences wide, branch angles mostly 20–40°; petals usually pale to bright yellow; pedicels and stems sometimes moderately septate-glandular.
var. austiniae
Source FNA vol. 9, p. 288. FNA vol. 9, p. 287.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis > Drymocallis lactea Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis
Sibling taxa
D. lactea var. austiniae
D. arguta, D. arizonica, D. ashlandica, D. campanulata, D. convallaria, D. cuneifolia, D. deseretica, D. fissa, D. glabrata, D. glandulosa, D. hansenii, D. micropetala, D. pseudorupestris, D. rhomboidea
Subordinate taxa
D. lactea var. austiniae, D. lactea var. lactea
Synonyms D. glandulosa subsp. nevadensis, Potentilla glandulosa subsp. nevadensis, P. glandulosa var. nevadensis Potentilla glandulosa var. lactea
Name authority unknown (Greene) Rydberg: in N. L. Britton et al., N. Amer. Fl. 22: 369. (1908)
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