Drymocallis hansenii
Drymocallis fissa
Hansen's cinquefoil, Hansen's drymocallis or wood beauty, Yosemite woodbeauty
bigflower cinquefoil, leafy drymocallis or wood beauty
short to elongate.
short to elongate.
usually solitary, sometimes loosely tufted, (3–)4–9 dm;
base (1–)2–4 mm diam., moderately to densely septate-glandular.
tufted to loosely spaced, (1.2–)1.5–3.5(–4.5) dm;
base (1.5–)2–3 mm diam., ± densely septate-glandular.
sparsely to moderately hairy;
basal (7–)10–25(–30) cm, leaflet pairs 3–4;
terminal leaflet broadly obovate, 2–5(–6) × 1.5–3.5(–4) cm, teeth single or double, 6–11 per side, apex usually rounded, sometimes obtuse;
cauline 1–3, proximally well developed, leaflet pairs 2–4.
sparsely to moderately hairy;
basal (3–)7–19 cm, leaflet pairs (4–)5–6(–10; additional reduced leaflets sometimes interspersed);
terminal leaflet usually broadly obovate-cuneate, sometimes elliptic, (1–)1.5–3.5(–5) × (1–)1.5–3(–3.5) cm, teeth single or double, 5–13 per side, apex rounded to obtuse;
cauline 1–3, well developed, leaflet pairs 4–6(–10).
10–20(–40)-flowered, not leafy, open, 1/4–2/3 of stem, narrow, branch angles 10–30°.
5–15-flowered, leafy, congested to ± open, 1/6–1/2 of stem, narrow to wide, branch angles 15–30(–40)°.
2–8 (proximal to 20) mm, predominantly short-hairy, sparsely to moderately septate-glandular.
1–12 mm, short-hairy, septate-glandular.
opening widely;
epicalyx bractlets linear to narrowly elliptic, 2–4 × 0.5–1 mm;
sepals spreading, 5–8 mm, apex acute to acuminate;
petals overlapping or not, spreading, cream-white to pale yellow, broadly obovate, 4–6 × 3–6 mm, usually longer than, sometimes equal to, sepals;
filaments 1.5–2.5(–3) mm, anthers 0.8–1 mm;
styles thickened, 0.8–1.2 mm.
opening widely;
epicalyx bractlets linear-oblanceolate, 3–7 × 1–2 mm;
sepals spreading, 6–10 mm, apex acute to acuminate;
petals overlapping, spreading, yellow, broadly obovate, 7–11 × 5–11 mm, equal to or exceeding sepals;
filaments 1.5–4.5 mm, anthers (0.7–)1–1.4 mm;
styles thickened, 1 mm.
light brown, 0.7–1 mm.
light brown, 1 mm.
= 14.
Drymocallis hansenii
Drymocallis fissa
D. D. Keck (in J. Clausen et al. 1940) speculated that Drymocallis hansenii was the stabilized recombinant of D. glandulosa var. reflexa and D. lactea. Alternatively, it may represent the California counterpart of D. convallaria, because it tends to have tall, thick-based, single stems and narrow inflorescences. The species is centered in the west-central Sierra Nevada of California, usually occurring in moist meadows and equivalent habitats. Plants near Lake Tahoe, which provide the high-elevation extreme, combine the smaller stature of D. lactea and the glandular-septate stem bases of D. hansenii; their optimal taxonomic disposition is uncertain.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Drymocallis fissa is distinctive in its relatively numerous leaflets (often with additional smaller ones), large flowers, and large, elongate anthers. It is most abundant in the Colorado Front Range, extending into the Medicine Bow Mountains of Wyoming. Outlying populations occur at least as far north as the Bighorn Mountains of northern Wyoming and the Black Hills of South Dakota. Tentatively included here are large-anthered populations from the eastern Uintah Mountains of Utah, though these often have fewer leaflets and smaller flowers of unknown color; they may represent a unique taxon worthy of separate recognition. Possible collections of D. fissa from New Mexico, including the type of Potentilla fissa var. major Torrey & A. Gray, are of uncertain placement in that they combine features of D. arguta, D. arizonica, and D. fissa. Reports from other states, including Montana, are probably all based on misidentified specimens.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
CA
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