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Hansen's cinquefoil, Hansen's drymocallis or wood beauty, Yosemite woodbeauty

Caudex branches

short to elongate.

elongate.

Stems

usually solitary, sometimes loosely tufted, (3–)4–9 dm;

base (1–)2–4 mm diam., moderately to densely septate-glandular.

openly tufted to loosely spaced, 0.5–4.5 dm;

base 1–2 mm diam., sparsely septate-glandular.

Leaves

sparsely to moderately hairy;

basal (7–)10–25(–30) cm, leaflet pairs 3–4;

terminal leaflet broadly obovate, 2–5(–6) × 1.5–3.5(–4) cm, teeth single or double, 6–11 per side, apex usually rounded, sometimes obtuse;

cauline 1–3, proximally well developed, leaflet pairs 2–4.

sparsely to moderately hairy;

basal 2–15 cm, leaflet pairs (2–)3–5;

terminal leaflet broadly cuneate to nearly round, 0.6–2(–3.5) × 0.5–2 cm, teeth irregularly single, 2–4 per side, apex rounded to truncate;

cauline 0–1(–2), reduced, leaflet pairs 1–2.

Inflorescences

10–20(–40)-flowered, not leafy, open, 1/4–2/3 of stem, narrow, branch angles 10–30°.

3–10(–15)-flowered, not leafy, open, 1/4–3/4 of stem, wide, branch angles 20–75°.

Pedicels

2–8 (proximal to 20) mm, predominantly short-hairy, sparsely to moderately septate-glandular.

2–15 (proximal to 30) mm, sparsely to moderately short-hairy, septate-glandular.

Flowers

opening widely;

epicalyx bractlets linear to narrowly elliptic, 2–4 × 0.5–1 mm;

sepals spreading, 5–8 mm, apex acute to acuminate;

petals overlapping or not, spreading, cream-white to pale yellow, broadly obovate, 4–6 × 3–6 mm, usually longer than, sometimes equal to, sepals;

filaments 1.5–2.5(–3) mm, anthers 0.8–1 mm;

styles thickened, 0.8–1.2 mm.

opening narrowly;

epicalyx bractlets linear to narrowly elliptic or lanceolate, 1–2 × 0.5 mm;

sepals erect, 2–5 mm, apex broadly obtuse;

petals scarcely overlapping, erect, yellow, narrowly obovate, 2–4 × 1.5–2.5 mm, ± equal to or slightly longer than sepals;

filaments 1.5–2.5 mm, anthers 0.5–0.8 mm;

styles slender, 1.5–2.5 mm.

Achenes

light brown, 0.7–1 mm.

brown, 0.8–1.2 mm.

2n

= 14.

Drymocallis hansenii

Drymocallis cuneifolia

Phenology Flowering Jun–Sep.
Habitat Moist ground, meadows, open forests, streamsides
Elevation 1200–1900(–2200) m (3900–6200(–7200) ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

D. D. Keck (in J. Clausen et al. 1940) speculated that Drymocallis hansenii was the stabilized recombinant of D. glandulosa var. reflexa and D. lactea. Alternatively, it may represent the California counterpart of D. convallaria, because it tends to have tall, thick-based, single stems and narrow inflorescences. The species is centered in the west-central Sierra Nevada of California, usually occurring in moist meadows and equivalent habitats. Plants near Lake Tahoe, which provide the high-elevation extreme, combine the smaller stature of D. lactea and the glandular-septate stem bases of D. hansenii; their optimal taxonomic disposition is uncertain.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Drymocallis cuneifolia has a complex nomenclatural history, complicated by the extreme rarity of the typical variety. Rydberg described the species on the basis of a single collection from the San Bernardino Mountains of San Bernardino County, probably near Green Lead Mines. When comparable (though significantly smaller) plants were found in the San Gabriel Mountains of Los Angeles County, P. A. Munz and I. M. Johnston (1925) adopted Potentilla cuneifolia (Rydberg) Th. Wolf for both extremes; this name was later replaced with P. peirsonii because of the earlier P. cuneifolia Bertoloni. In the absence of comparable new collections from the San Bernardino Mountains, D. D. Keck (in J. Clausen et al. 1940) concluded that the type of D. cuneifolia was merely an immature specimen of D. lactea var. lactea and accordingly described P. glandulosa subsp. ewanii to accommodate populations in the San Gabriel Mountains.

The 2004 discovery of a small population of plants comparable to the type of Drymocallis cuneifolia from near the type locality (Elvin 3555, IRVC, UCR) confirms that D. cuneifolia is a valid taxon that shares most of the diagnostic features of populations in the San Gabriel Mountains. Plants in the San Bernardino Mountains tend to be larger than those in the San Gabriel Mountains, with fewer and somewhat differently shaped leaflets; they are accordingly treated here as varieties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems 2–4.5 dm; basal leaves 5–15 cm; San Bernardino Mountains.
var. cuneifolia
1. Stems 0.5–2(–2.5) dm; basal leaves 2–10 cm; San Gabriel Mountains.
var. ewanii
Source FNA vol. 9, p. 287. FNA vol. 9, p. 294.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis
Sibling taxa
D. arguta, D. arizonica, D. ashlandica, D. campanulata, D. convallaria, D. cuneifolia, D. deseretica, D. fissa, D. glabrata, D. glandulosa, D. lactea, D. micropetala, D. pseudorupestris, D. rhomboidea
D. arguta, D. arizonica, D. ashlandica, D. campanulata, D. convallaria, D. deseretica, D. fissa, D. glabrata, D. glandulosa, D. hansenii, D. lactea, D. micropetala, D. pseudorupestris, D. rhomboidea
Subordinate taxa
D. cuneifolia var. cuneifolia, D. cuneifolia var. ewanii
Synonyms Potentilla hansenii, D. glandulosa subsp. hansenii, P. glandulosa subsp. hansenii Potentilla peirsonii
Name authority (Greene) Rydberg: Monogr. N. Amer. Potentilleae, 200. (1898) Rydberg: Monogr. N. Amer. Potentilleae, 204, plate 111 [as cuneata]. (1898)
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