FNA: Drymocallis hansenii vs. Drymocallis ashlandica

	Drymocallis hansenii	Drymocallis ashlandica
	Hansen's cinquefoil, Hansen's drymocallis or wood beauty, Yosemite woodbeauty	Ashland cinquefoil, Mazama drymocallis or wood beauty
Caudex branches	short to elongate.	short.
Stems	usually solitary, sometimes loosely tufted, (3–)4–9 dm; base (1–)2–4 mm diam., moderately to densely septate-glandular.	tufted, (1–)2–5 dm; base 2–3 mm diam., ± densely septate-glandular.
Leaves	sparsely to moderately hairy; basal (7–)10–25(–30) cm, leaflet pairs 3–4; terminal leaflet broadly obovate, 2–5(–6) × 1.5–3.5(–4) cm, teeth single or double, 6–11 per side, apex usually rounded, sometimes obtuse; cauline 1–3, proximally well developed, leaflet pairs 2–4.	moderately to ± densely hairy; basal 6–16 cm, leaflet pairs 2–3(–4); terminal leaflet broadly obovate, 1.5–4 × 1–3 cm, teeth ± double, 7–12 per side, apex rounded; cauline 0–2, moderately developed, leaflet pairs 2–3.
Inflorescences	10–20(–40)-flowered, not leafy, open, 1/4–2/3 of stem, narrow, branch angles 10–30°.	5–15(–20)-flowered, not leafy, congested to deeply branched, 1/10–2/3 or less of stem, usually narrow, rarely ± wide, branch angles (5–)10–20(–30)°.
Pedicels	2–8 (proximal to 20) mm, predominantly short-hairy, sparsely to moderately septate-glandular.	2–10 (proximal to 30) mm, sparsely to moderately short-hairy, predominantly septate-glandular, often obscuring other vestiture.
Flowers	opening widely; epicalyx bractlets linear to narrowly elliptic, 2–4 × 0.5–1 mm; sepals spreading, 5–8 mm, apex acute to acuminate; petals overlapping or not, spreading, cream-white to pale yellow, broadly obovate, 4–6 × 3–6 mm, usually longer than, sometimes equal to, sepals; filaments 1.5–2.5(–3) mm, anthers 0.8–1 mm; styles thickened, 0.8–1.2 mm.	opening widely; epicalyx bractlets linear to broadly elliptic, 3–5 × 1–2 mm; sepals spreading, 5–7(–9) mm, apex acute; petals overlapping, spreading, light yellow, broadly obovate, 5–9 × 4–7 mm, equal to or longer than sepals; filaments 1.5–5 mm, anthers 0.7–1.2 mm; styles ± thickened, 1 mm.
Achenes	light brown, 0.7–1 mm.	light brown, 1 mm.
2n	= 14.

	Drymocallis hansenii	Drymocallis ashlandica
Phenology	Flowering Jun–Sep.	Flowering Jun–Aug.
Habitat	Moist ground, meadows, open forests, streamsides	Meadows, roadsides, sandy shorelines
Elevation	1200–1900(–2200) m (3900–6200(–7200) ft)	500–2000 m (1600–6600 ft)
Distribution	CA [WildflowerSearch map] [BONAP county map]	OR [WildflowerSearch map] [BONAP county map]
Discussion	D. D. Keck (in J. Clausen et al. 1940) speculated that Drymocallis hansenii was the stabilized recombinant of D. glandulosa var. reflexa and D. lactea. Alternatively, it may represent the California counterpart of D. convallaria, because it tends to have tall, thick-based, single stems and narrow inflorescences. The species is centered in the west-central Sierra Nevada of California, usually occurring in moist meadows and equivalent habitats. Plants near Lake Tahoe, which provide the high-elevation extreme, combine the smaller stature of D. lactea and the glandular-septate stem bases of D. hansenii; their optimal taxonomic disposition is uncertain. (Discussion copyrighted by Flora of North America; reprinted with permission.)	The name Potentilla glandulosa subsp. ashlandica was misapplied to Drymocallis lactea var. austiniae (B. Ertter 1993); as circumscribed here, D. ashlandica is a poorly known species restricted to the Cascade and Siskiyou ranges of southwestern Oregon. It very likely also occurs in adjacent California; no collections from there are known. Drymocallis ashlandica differs from most sympatric populations of D. lactea var. austiniae and resembles D. pseudorupestris in its densely glandular inflorescences, which tend to be more densely congested and/or narrower than those of the other two species. Drymocallis ashlandica was described by Greene as Potentilla ashlandica, and by Howell as P. ciliata (a later homonym), both based on a collection by Howell from the Siskiyou Mountains near Ashland, Oregon. The circumscription here, which matches that of D. D. Keck (in J. Clausen et al. 1940) and M. E. Peck (1961), encompasses populations from near Oregon Caves and Ashland in the Siskiyou Mountains to near McKenzie Bridge, Lane County, and Sparks Lake, Deschutes County. It is currently known from relatively few collections, mostly from the Crater Lake area, and may prove to be of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 9, p. 287.	FNA vol. 9, p. 288.
Parent taxa	Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis	Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis
Sibling taxa	D. arguta, D. arizonica, D. ashlandica, D. campanulata, D. convallaria, D. cuneifolia, D. deseretica, D. fissa, D. glabrata, D. glandulosa, D. lactea, D. micropetala, D. pseudorupestris, D. rhomboidea	D. arguta, D. arizonica, D. campanulata, D. convallaria, D. cuneifolia, D. deseretica, D. fissa, D. glabrata, D. glandulosa, D. hansenii, D. lactea, D. micropetala, D. pseudorupestris, D. rhomboidea
Synonyms	Potentilla hansenii, D. glandulosa subsp. hansenii, P. glandulosa subsp. hansenii	Potentilla ashlandica, D. glandulosa subsp. ashlandica, P. glandulosa subsp. ashlandica
Name authority	(Greene) Rydberg: Monogr. N. Amer. Potentilleae, 200. (1898)	(Greene) Rydberg: Monogr. N. Amer. Potentilleae, 200. (1898)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: OR Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Drymocallis ashlandica

Hansen's cinquefoil, Hansen's drymocallis or wood beauty, Yosemite woodbeauty

Ashland cinquefoil, Mazama drymocallis or wood beauty

Caudex branches

short to elongate.

short.

Stems

usually solitary, sometimes loosely tufted, (3–)4–9 dm;

base (1–)2–4 mm diam., moderately to densely septate-glandular.

tufted, (1–)2–5 dm;

base 2–3 mm diam., ± densely septate-glandular.

Leaves

sparsely to moderately hairy;

basal (7–)10–25(–30) cm, leaflet pairs 3–4;

terminal leaflet broadly obovate, 2–5(–6) × 1.5–3.5(–4) cm, teeth single or double, 6–11 per side, apex usually rounded, sometimes obtuse;

cauline 1–3, proximally well developed, leaflet pairs 2–4.

moderately to ± densely hairy;

basal 6–16 cm, leaflet pairs 2–3(–4);

terminal leaflet broadly obovate, 1.5–4 × 1–3 cm, teeth ± double, 7–12 per side, apex rounded;

cauline 0–2, moderately developed, leaflet pairs 2–3.

Inflorescences

10–20(–40)-flowered, not leafy, open, 1/4–2/3 of stem, narrow, branch angles 10–30°.

5–15(–20)-flowered, not leafy, congested to deeply branched, 1/10–2/3 or less of stem, usually narrow, rarely ± wide, branch angles (5–)10–20(–30)°.

Pedicels

2–8 (proximal to 20) mm, predominantly short-hairy, sparsely to moderately septate-glandular.

2–10 (proximal to 30) mm, sparsely to moderately short-hairy, predominantly septate-glandular, often obscuring other vestiture.

Flowers

opening widely;

epicalyx bractlets linear to narrowly elliptic, 2–4 × 0.5–1 mm;

sepals spreading, 5–8 mm, apex acute to acuminate;

petals overlapping or not, spreading, cream-white to pale yellow, broadly obovate, 4–6 × 3–6 mm, usually longer than, sometimes equal to, sepals;

filaments 1.5–2.5(–3) mm, anthers 0.8–1 mm;

styles thickened, 0.8–1.2 mm.

opening widely;

epicalyx bractlets linear to broadly elliptic, 3–5 × 1–2 mm;

sepals spreading, 5–7(–9) mm, apex acute;

petals overlapping, spreading, light yellow, broadly obovate, 5–9 × 4–7 mm, equal to or longer than sepals;

filaments 1.5–5 mm, anthers 0.7–1.2 mm;

styles ± thickened, 1 mm.

Achenes

light brown, 0.7–1 mm.

light brown, 1 mm.

= 14.

Drymocallis ashlandica

Phenology

Flowering Jun–Sep.

Flowering Jun–Aug.

Habitat

Moist ground, meadows, open forests, streamsides

Meadows, roadsides, sandy shorelines

Elevation

1200–1900(–2200) m (3900–6200(–7200) ft)

500–2000 m (1600–6600 ft)

Distribution

[WildflowerSearch map]

[BONAP county map]

[WildflowerSearch map]

[BONAP county map]

Discussion

D. D. Keck (in J. Clausen et al. 1940) speculated that Drymocallis hansenii was the stabilized recombinant of D. glandulosa var. reflexa and D. lactea. Alternatively, it may represent the California counterpart of D. convallaria, because it tends to have tall, thick-based, single stems and narrow inflorescences. The species is centered in the west-central Sierra Nevada of California, usually occurring in moist meadows and equivalent habitats. Plants near Lake Tahoe, which provide the high-elevation extreme, combine the smaller stature of D. lactea and the glandular-septate stem bases of D. hansenii; their optimal taxonomic disposition is uncertain.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The name Potentilla glandulosa subsp. ashlandica was misapplied to Drymocallis lactea var. austiniae (B. Ertter 1993); as circumscribed here, D. ashlandica is a poorly known species restricted to the Cascade and Siskiyou ranges of southwestern Oregon. It very likely also occurs in adjacent California; no collections from there are known. Drymocallis ashlandica differs from most sympatric populations of D. lactea var. austiniae and resembles D. pseudorupestris in its densely glandular inflorescences, which tend to be more densely congested and/or narrower than those of the other two species.

Drymocallis ashlandica was described by Greene as Potentilla ashlandica, and by Howell as P. ciliata (a later homonym), both based on a collection by Howell from the Siskiyou Mountains near Ashland, Oregon. The circumscription here, which matches that of D. D. Keck (in J. Clausen et al. 1940) and M. E. Peck (1961), encompasses populations from near Oregon Caves and Ashland in the Siskiyou Mountains to near McKenzie Bridge, Lane County, and Sparks Lake, Deschutes County. It is currently known from relatively few collections, mostly from the Crater Lake area, and may prove to be of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 9, p. 287.

FNA vol. 9, p. 288.

Parent taxa

Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis

Sibling taxa

D. arguta, D. arizonica, D. ashlandica, D. campanulata, D. convallaria, D. cuneifolia, D. deseretica, D. fissa, D. glabrata, D. glandulosa, D. lactea, D. micropetala, D. pseudorupestris, D. rhomboidea

D. arguta, D. arizonica, D. campanulata, D. convallaria, D. cuneifolia, D. deseretica, D. fissa, D. glabrata, D. glandulosa, D. hansenii, D. lactea, D. micropetala, D. pseudorupestris, D. rhomboidea

Synonyms

Potentilla hansenii, D. glandulosa subsp. hansenii, P. glandulosa subsp. hansenii

Potentilla ashlandica, D. glandulosa subsp. ashlandica, P. glandulosa subsp. ashlandica

Name authority

(Greene) Rydberg: Monogr. N. Amer. Potentilleae, 200. (1898)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora

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Drymocallis hansenii

Drymocallis ashlandica

Drymocallis hansenii

Drymocallis ashlandica