Drymocallis hansenii |
Drymocallis arguta |
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Hansen's cinquefoil, Hansen's drymocallis or wood beauty, Yosemite woodbeauty |
cordilleran drymocallis, sticky cinquefoil, tall cinquefoil, tall drymocallis or wood beauty, tall wood-beauty |
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Caudex branches | short to elongate. |
short. |
Stems | usually solitary, sometimes loosely tufted, (3–)4–9 dm; base (1–)2–4 mm diam., moderately to densely septate-glandular. |
usually solitary, sometimes tufted, (1–)3–10 dm; base 3–6 mm diam., ± densely, sometimes sparsely, septate-glandular. |
Leaves | sparsely to moderately hairy; basal (7–)10–25(–30) cm, leaflet pairs 3–4; terminal leaflet broadly obovate, 2–5(–6) × 1.5–3.5(–4) cm, teeth single or double, 6–11 per side, apex usually rounded, sometimes obtuse; cauline 1–3, proximally well developed, leaflet pairs 2–4. |
usually moderately to densely hairy; basal (6–)12–40 cm, leaflet pairs (3–)4–5; terminal leaflet broadly elliptic-obovate, (2–)4–10 × (1–)2–4.5 cm, teeth mostly double, 15–30+ per side, apex obtuse to acute; cauline 2–3, relatively well developed, leaflet pairs 2–4. |
Inflorescences | 10–20(–40)-flowered, not leafy, open, 1/4–2/3 of stem, narrow, branch angles 10–30°. |
10–40-flowered, not leafy, congested, sometimes more elongate in fruit, 1/10–1/5(–1/3) of stem, narrow, branch angles 5–20°. |
Pedicels | 2–8 (proximal to 20) mm, predominantly short-hairy, sparsely to moderately septate-glandular. |
1–5 (proximal to 15) mm, densely short-hairy, septate-glandular. |
Flowers | opening widely; epicalyx bractlets linear to narrowly elliptic, 2–4 × 0.5–1 mm; sepals spreading, 5–8 mm, apex acute to acuminate; petals overlapping or not, spreading, cream-white to pale yellow, broadly obovate, 4–6 × 3–6 mm, usually longer than, sometimes equal to, sepals; filaments 1.5–2.5(–3) mm, anthers 0.8–1 mm; styles thickened, 0.8–1.2 mm. |
opening widely; epicalyx bractlets ± elliptic, 4–6(–8) × 1–2 mm; sepals spreading, (5–)7–10 mm, apex obtuse to acute, apiculate; petals overlapping, spreading, cream-white to yellowish, broadly elliptic, (5–)7–9 × (5–)6–8 mm, ± equal to or slightly longer than sepals; filaments 1.5–2.5 mm, anthers 0.7–1 mm; styles thickened, 1 mm. |
Achenes | light brown, 0.7–1 mm. |
light brown, 1 mm. |
2n | = 14. |
= 14. |
Drymocallis hansenii |
Drymocallis arguta |
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Phenology | Flowering Jun–Sep. | Flowering May–Aug. |
Habitat | Moist ground, meadows, open forests, streamsides | Prairies, abandoned pastures, open woodlands, shallow soil and grassy openings on ridges, slopes, bluffs, and rocky barrens |
Elevation | 1200–1900(–2200) m (3900–6200(–7200) ft) | 10–2300 m (0–7500 ft) |
Distribution |
CA
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AR; CO; CT; IA; IL; IN; KS; MA; MD; ME; MI; MN; MO; MT; ND; NE; NH; NJ; NM; NY; OH; OK; PA; SD; TN; VA; VT; WI; WV; WY; AB; MB; NB; NT; ON; QC; SK; YT
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Discussion | D. D. Keck (in J. Clausen et al. 1940) speculated that Drymocallis hansenii was the stabilized recombinant of D. glandulosa var. reflexa and D. lactea. Alternatively, it may represent the California counterpart of D. convallaria, because it tends to have tall, thick-based, single stems and narrow inflorescences. The species is centered in the west-central Sierra Nevada of California, usually occurring in moist meadows and equivalent habitats. Plants near Lake Tahoe, which provide the high-elevation extreme, combine the smaller stature of D. lactea and the glandular-septate stem bases of D. hansenii; their optimal taxonomic disposition is uncertain. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Drymocallis arguta is the only species of the genus occurring east of the Rocky Mountains, except for D. fissa in immediately adjacent prairies and the Black Hills of South Dakota. It is primarily a species of intact prairies and pasturelands throughout the Great Plains, extending as scattered populations in equivalent habitats throughout the northeastern states and adjacent provinces. Some of these states consider it a species of conservation concern (as Potentilla arguta, tall cinquefoil), and the sparsely hairy nature of some of these populations suggests that further taxonomic attention might be warranted. Drymocallis arguta occurs also in the Colorado Front Range, generally at elevations higher than D. fissa, with which it sometimes intergrades morphologically. It intergrades also with D. convallaria but tends to be larger, coarser, and more densely hairy with more acute leaflets. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 287. | FNA vol. 9, p. 285. |
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis |
Sibling taxa | ||
Synonyms | Potentilla hansenii, D. glandulosa subsp. hansenii, P. glandulosa subsp. hansenii | Potentilla arguta, D. agrimonoides |
Name authority | (Greene) Rydberg: Monogr. N. Amer. Potentilleae, 200. (1898) | (Pursh) Rydberg: Monogr. N. Amer. Potentilleae, 192. (1898) |
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