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Idaho drymocallis, Idaho drymocallis or wood beauty

Caudex branches

short.

elongate.

Stems

± tufted, (2–)2.5–6(–8) dm;

base 2–3 mm diam., moderately to densely septate-glandular.

openly tufted to loosely spaced, 0.5–4.5 dm;

base 1–2 mm diam., sparsely septate-glandular.

Leaves

glabrate or sparsely hairy (hairier at northern margin of range);

basal (8–)20–30 cm, leaflet pairs 2–4;

terminal leaflet ± obovate, 2–6(–7) × (1–)1.5–3 cm, teeth ± double, 5–25 per side, apex obtuse to acute;

cauline 0–2, well developed, leaflet pairs 1–3.

sparsely to moderately hairy;

basal 2–15 cm, leaflet pairs (2–)3–5;

terminal leaflet broadly cuneate to nearly round, 0.6–2(–3.5) × 0.5–2 cm, teeth irregularly single, 2–4 per side, apex rounded to truncate;

cauline 0–1(–2), reduced, leaflet pairs 1–2.

Inflorescences

(5–)10–30(–60)-flowered, leafy, open, 1/6–2/3 of stem, wide, branch angles (20–)30–60°.

3–10(–15)-flowered, not leafy, open, 1/4–3/4 of stem, wide, branch angles 20–75°.

Pedicels

5–40 (proximal to 45) mm, sparsely to moderately short-hairy, predominantly septate-glandular.

2–15 (proximal to 30) mm, sparsely to moderately short-hairy, septate-glandular.

Flowers

opening widely;

epicalyx bractlets linear-oblanceolate, 2–6 × 0.5–1 mm;

sepals spreading, 4–7(–8) mm, apex usually acute, rarely obtuse;

petals ± overlapping, spreading, usually bright, sometimes pale, yellow, broadly obovate to round, 5–12 × (4–)5–11 mm, usually longer, rarely shorter, than sepals;

filaments 1.5–4 mm, anthers 0.7–1 mm;

styles thickened, 1 mm.

opening narrowly;

epicalyx bractlets linear to narrowly elliptic or lanceolate, 1–2 × 0.5 mm;

sepals erect, 2–5 mm, apex broadly obtuse;

petals scarcely overlapping, erect, yellow, narrowly obovate, 2–4 × 1.5–2.5 mm, ± equal to or slightly longer than sepals;

filaments 1.5–2.5 mm, anthers 0.5–0.8 mm;

styles slender, 1.5–2.5 mm.

Achenes

brown to reddish, 1.2 mm.

brown, 0.8–1.2 mm.

Drymocallis glabrata

Drymocallis cuneifolia

Phenology Flowering May–Aug.
Habitat Montane meadows, open forests
Elevation 400–2800 m (1300–9200 ft)
Distribution
from FNA
ID; MT; NV; OR; UT; WA; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Drymocallis glabrata is the common member of the genus in central Idaho and adjacent parts of neighboring states, characterized by acute, glabrate leaves, leafy, widely branched inflorescences, relatively long pedicels with slender, septate glands, linear-oblanceolate epicalyx bractlets, and relatively large, bright yellow petals. Plants from northern Idaho, at the lower elevational extreme, tend to be hairier than those elsewhere in the species range. The species enters Utah in Cache and Weber counties; most other Utah populations placed by S. L. Welsh et al. (1993) in this taxon (as Potentilla glandulosa var. intermedia) are treated here as D. deseretica.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Drymocallis cuneifolia has a complex nomenclatural history, complicated by the extreme rarity of the typical variety. Rydberg described the species on the basis of a single collection from the San Bernardino Mountains of San Bernardino County, probably near Green Lead Mines. When comparable (though significantly smaller) plants were found in the San Gabriel Mountains of Los Angeles County, P. A. Munz and I. M. Johnston (1925) adopted Potentilla cuneifolia (Rydberg) Th. Wolf for both extremes; this name was later replaced with P. peirsonii because of the earlier P. cuneifolia Bertoloni. In the absence of comparable new collections from the San Bernardino Mountains, D. D. Keck (in J. Clausen et al. 1940) concluded that the type of D. cuneifolia was merely an immature specimen of D. lactea var. lactea and accordingly described P. glandulosa subsp. ewanii to accommodate populations in the San Gabriel Mountains.

The 2004 discovery of a small population of plants comparable to the type of Drymocallis cuneifolia from near the type locality (Elvin 3555, IRVC, UCR) confirms that D. cuneifolia is a valid taxon that shares most of the diagnostic features of populations in the San Gabriel Mountains. Plants in the San Bernardino Mountains tend to be larger than those in the San Gabriel Mountains, with fewer and somewhat differently shaped leaflets; they are accordingly treated here as varieties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems 2–4.5 dm; basal leaves 5–15 cm; San Bernardino Mountains.
var. cuneifolia
1. Stems 0.5–2(–2.5) dm; basal leaves 2–10 cm; San Gabriel Mountains.
var. ewanii
Source FNA vol. 9, p. 291. FNA vol. 9, p. 294.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis
Sibling taxa
D. arguta, D. arizonica, D. ashlandica, D. campanulata, D. convallaria, D. cuneifolia, D. deseretica, D. fissa, D. glandulosa, D. hansenii, D. lactea, D. micropetala, D. pseudorupestris, D. rhomboidea
D. arguta, D. arizonica, D. ashlandica, D. campanulata, D. convallaria, D. deseretica, D. fissa, D. glabrata, D. glandulosa, D. hansenii, D. lactea, D. micropetala, D. pseudorupestris, D. rhomboidea
Subordinate taxa
D. cuneifolia var. cuneifolia, D. cuneifolia var. ewanii
Synonyms D. glandulosa subsp. glabrata, Potentilla glandulosa subsp. glabrata, P. glandulosa var. incisa, P. glandulosa var. intermedia Potentilla peirsonii
Name authority Rydberg: Monogr. N. Amer. Potentilleae, 201, plate 109. (1898) Rydberg: Monogr. N. Amer. Potentilleae, 204, plate 111 [as cuneata]. (1898)
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