Drymocallis fissa
Drymocallis campanulata
bigflower cinquefoil, leafy drymocallis or wood beauty
John Day drymocallis, John Day drymocallis or wood beauty
short to elongate.
elongate.
tufted to loosely spaced, (1.2–)1.5–3.5(–4.5) dm;
base (1.5–)2–3 mm diam., ± densely septate-glandular.
openly tufted to loosely spaced, 1.5–4.5 dm;
base 1.2–3.5 mm diam., ± densely septate-glandular.
sparsely to moderately hairy;
basal (3–)7–19 cm, leaflet pairs (4–)5–6(–10; additional reduced leaflets sometimes interspersed);
terminal leaflet usually broadly obovate-cuneate, sometimes elliptic, (1–)1.5–3.5(–5) × (1–)1.5–3(–3.5) cm, teeth single or double, 5–13 per side, apex rounded to obtuse;
cauline 1–3, well developed, leaflet pairs 4–6(–10).
sparsely to ± densely hairy;
basal 6–20 cm, leaflet pairs (2–)3–4(–5);
terminal leaflet broadly obovate to nearly round, 1–4 × 1–3.5 cm, teeth usually ± double, 5–10 per side, apex rounded;
cauline 0–2, well developed, leaflet pairs 2–3.
5–15-flowered, leafy, congested to ± open, 1/6–1/2 of stem, narrow to wide, branch angles 15–30(–40)°.
5–50-flowered, leafy, open, (1/5–)1/2(–4/5) of stem, wide, branch angles 25–50°.
1–12 mm, short-hairy, septate-glandular.
2–5 (proximal to 10) mm, sparsely to moderately short-hairy, predominantly septate-glandular.
opening widely;
epicalyx bractlets linear-oblanceolate, 3–7 × 1–2 mm;
sepals spreading, 6–10 mm, apex acute to acuminate;
petals overlapping, spreading, yellow, broadly obovate, 7–11 × 5–11 mm, equal to or exceeding sepals;
filaments 1.5–4.5 mm, anthers (0.7–)1–1.4 mm;
styles thickened, 1 mm.
opening narrowly;
epicalyx bractlets usually lanceolate, sometimes linear, 3–4 × 1–1.5 mm;
sepals ± erect, 5–8 mm, apex acute to obtuse;
petals overlapping, ± erect, light yellow, broadly obovate, 5–11 × (3–)5–8 mm, equal to or exceeding sepals;
filaments 1.5–3 mm, anthers 1 mm;
styles slender, (1–)1.5–2.5 mm.
light brown, 1 mm.
light brown, 1 mm.
Drymocallis fissa
Drymocallis campanulata
Drymocallis fissa is distinctive in its relatively numerous leaflets (often with additional smaller ones), large flowers, and large, elongate anthers. It is most abundant in the Colorado Front Range, extending into the Medicine Bow Mountains of Wyoming. Outlying populations occur at least as far north as the Bighorn Mountains of northern Wyoming and the Black Hills of South Dakota. Tentatively included here are large-anthered populations from the eastern Uintah Mountains of Utah, though these often have fewer leaflets and smaller flowers of unknown color; they may represent a unique taxon worthy of separate recognition. Possible collections of D. fissa from New Mexico, including the type of Potentilla fissa var. major Torrey & A. Gray, are of uncertain placement in that they combine features of D. arguta, D. arizonica, and D. fissa. Reports from other states, including Montana, are probably all based on misidentified specimens.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Drymocallis campanulata is one of the more distinct species in the genus, with its campanulate flowers, relatively large butter yellow petals, and densely glandular pedicels. It occurs primarily in the canyons of the John Day River in Grant and Wheeler counties.
The invalidly published Potentilla campanulata D. D. Keck provides a full description and illustration for the subsequent basionym.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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