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John Day drymocallis, John Day drymocallis or wood beauty

Drymocallis pseudorupestris

cliff drymocallis, cliff woodbeauty, false rock loving cinquefoil, Rocky Mountain sticky cinquefoil

Caudex branches

elongate.

elongate.

Stems

openly tufted to loosely spaced, 1.5–4.5 dm;

base 1.2–3.5 mm diam., ± densely septate-glandular.

openly tufted to loosely spaced, (0.3–)0.6–4 dm;

base 1–3 mm diam., ± densely septate-glandular.

Leaves

sparsely to ± densely hairy;

basal 6–20 cm, leaflet pairs (2–)3–4(–5);

terminal leaflet broadly obovate to nearly round, 1–4 × 1–3.5 cm, teeth usually ± double, 5–10 per side, apex rounded;

cauline 0–2, well developed, leaflet pairs 2–3.

glabrate or sparsely to ± densely hairy;

basal (2–)3–16 cm, leaflet pairs (2–)3–4(–5);

terminal leaflet broadly obovate-cuneate to flabellate, 0.2–3(–4) × 0.5–3 cm, teeth single or double, 2–15 per side, apex usually rounded to truncate, sometimes obtuse;

cauline 0–2, reduced, leaflet pairs 2–3.

Inflorescences

5–50-flowered, leafy, open, (1/5–)1/2(–4/5) of stem, wide, branch angles 25–50°.

2–40-flowered, not or ± leafy, open, 1/6–3/4(–4/5) of stem, ± wide, branch angles (10–)20–40(–50)°.

Pedicels

2–5 (proximal to 10) mm, sparsely to moderately short-hairy, predominantly septate-glandular.

3–20 (proximal to 40) mm, not or sparsely to moderately short-hairy, predominantly septate-glandular.

Flowers

opening narrowly;

epicalyx bractlets usually lanceolate, sometimes linear, 3–4 × 1–1.5 mm;

sepals ± erect, 5–8 mm, apex acute to obtuse;

petals overlapping, ± erect, light yellow, broadly obovate, 5–11 × (3–)5–8 mm, equal to or exceeding sepals;

filaments 1.5–3 mm, anthers 1 mm;

styles slender, (1–)1.5–2.5 mm.

opening widely;

epicalyx bractlets linear to elliptic, 2–6 × 1–2 mm;

sepals spreading, 4–7(–9) mm, apex acute to obtuse, apiculate;

petals overlapping or not, spreading, cream-white to pale yellow (red-tinged in var. crumiana), narrowly to broadly obovate, 4–12 × 3–11 mm, longer than sepals;

filaments 1–4 mm, anthers 0.7–1.2 mm;

styles thickened, 1–1.5 mm.

Achenes

light brown, 1 mm.

light brown, 1 mm.

Drymocallis campanulata

Drymocallis pseudorupestris

Phenology Flowering May–Jul.
Habitat Basaltic cliffs and talus above streams
Elevation 700–1400 m (2300–4600 ft)
Distribution
from FNA
OR
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; ID; MT; NV; OR; UT; WA; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Drymocallis campanulata is one of the more distinct species in the genus, with its campanulate flowers, relatively large butter yellow petals, and densely glandular pedicels. It occurs primarily in the canyons of the John Day River in Grant and Wheeler counties.

The invalidly published Potentilla campanulata D. D. Keck provides a full description and illustration for the subsequent basionym.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

Drymocallis pseudorupestris occurs from Alberta and Washington to California and Utah, mostly in montane habitats; it is the species most often associated with rocky habitats, including talus slopes, for which its relatively elongate caudex branches are an obvious adaptation. Vestiture is dominated by abundant septate glands on stems and in the inflorescences. Except for var. pseudorupestris, which occurs only in the northeastern part of the species range, plants are relatively short, usually less than 2.5 dm. Three intergrading varieties accommodate the extremes at the northeastern and southern ends of the range.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems (1–)2–4 dm, bases (1.5–)2–3 mm diam.; basal leaves (4–)7–16 cm; terminal leaflets (1–)2–3(–4) cm, teeth usually double, (5–)8–15 per side; flowers (5–)10–40; petals 6–12 × 5–11 mm, widely overlapping; filaments 2–4 mm.
var. pseudorupestris
1. Stems (0.3–)0.6–2.5 dm, bases 1–2(–3) mm diam.; basal leaves (2–)3–9(–15) cm; terminal leaflets 0.2–2(–4) cm, teeth single or ± double, 2–8(–12) per side; flowers 2–12(–20); petals 4–8(–9) × 3–6(–8) mm, not or ± overlapping; filaments 1–2.5(–3) mm
→ 2
2. Basal leaves: leaflet pairs (2–)3(–4); hypanthia and sepals not bristly or bristles less than 1 mm; short hairs sparse to moderately abundant on stems and pedicels (sometimes absent); styles usually golden brown, rarely reddish.
var. saxicola
2. Basal leaves: leaflet pairs 3–4(–5); hypanthia and sepals prominently bristly, bristles 1–1.5 mm; short hairs absent or sparse on stems and pedicels; styles usually dark red, rarely golden brown.
var. crumiana
Source FNA vol. 9, p. 294. FNA vol. 9, p. 289.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis
Sibling taxa
D. arguta, D. arizonica, D. ashlandica, D. convallaria, D. cuneifolia, D. deseretica, D. fissa, D. glabrata, D. glandulosa, D. hansenii, D. lactea, D. micropetala, D. pseudorupestris, D. rhomboidea
D. arguta, D. arizonica, D. ashlandica, D. campanulata, D. convallaria, D. cuneifolia, D. deseretica, D. fissa, D. glabrata, D. glandulosa, D. hansenii, D. lactea, D. micropetala, D. rhomboidea
Subordinate taxa
D. pseudorupestris var. crumiana, D. pseudorupestris var. pseudorupestris, D. pseudorupestris var. saxicola
Synonyms Potentilla glandulosa var. campanulata Potentilla pseudorupestris, D. glandulosa subsp. pseudorupestris, P. glandulosa subsp. pseudorupestris, P. glandulosa var. pseudorupestris, P. rupestris var. americana
Name authority (C. L. Hitchcock) Ertter: J. Bot. Res. Inst. Texas 1: 43. (2007) (Rydberg) Rydberg: Monogr. N. Amer. Potentilleae, 194. (1898)
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