Drymocallis campanulata |
Drymocallis pseudorupestris |
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John Day drymocallis, John Day drymocallis or wood beauty |
cliff drymocallis, cliff woodbeauty, false rock loving cinquefoil, Rocky Mountain sticky cinquefoil |
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Caudex branches | elongate. |
elongate. |
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Stems | openly tufted to loosely spaced, 1.5–4.5 dm; base 1.2–3.5 mm diam., ± densely septate-glandular. |
openly tufted to loosely spaced, (0.3–)0.6–4 dm; base 1–3 mm diam., ± densely septate-glandular. |
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Leaves | sparsely to ± densely hairy; basal 6–20 cm, leaflet pairs (2–)3–4(–5); terminal leaflet broadly obovate to nearly round, 1–4 × 1–3.5 cm, teeth usually ± double, 5–10 per side, apex rounded; cauline 0–2, well developed, leaflet pairs 2–3. |
glabrate or sparsely to ± densely hairy; basal (2–)3–16 cm, leaflet pairs (2–)3–4(–5); terminal leaflet broadly obovate-cuneate to flabellate, 0.2–3(–4) × 0.5–3 cm, teeth single or double, 2–15 per side, apex usually rounded to truncate, sometimes obtuse; cauline 0–2, reduced, leaflet pairs 2–3. |
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Inflorescences | 5–50-flowered, leafy, open, (1/5–)1/2(–4/5) of stem, wide, branch angles 25–50°. |
2–40-flowered, not or ± leafy, open, 1/6–3/4(–4/5) of stem, ± wide, branch angles (10–)20–40(–50)°. |
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Pedicels | 2–5 (proximal to 10) mm, sparsely to moderately short-hairy, predominantly septate-glandular. |
3–20 (proximal to 40) mm, not or sparsely to moderately short-hairy, predominantly septate-glandular. |
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Flowers | opening narrowly; epicalyx bractlets usually lanceolate, sometimes linear, 3–4 × 1–1.5 mm; sepals ± erect, 5–8 mm, apex acute to obtuse; petals overlapping, ± erect, light yellow, broadly obovate, 5–11 × (3–)5–8 mm, equal to or exceeding sepals; filaments 1.5–3 mm, anthers 1 mm; styles slender, (1–)1.5–2.5 mm. |
opening widely; epicalyx bractlets linear to elliptic, 2–6 × 1–2 mm; sepals spreading, 4–7(–9) mm, apex acute to obtuse, apiculate; petals overlapping or not, spreading, cream-white to pale yellow (red-tinged in var. crumiana), narrowly to broadly obovate, 4–12 × 3–11 mm, longer than sepals; filaments 1–4 mm, anthers 0.7–1.2 mm; styles thickened, 1–1.5 mm. |
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Achenes | light brown, 1 mm. |
light brown, 1 mm. |
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Drymocallis campanulata |
Drymocallis pseudorupestris |
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Phenology | Flowering May–Jul. | |||||||||
Habitat | Basaltic cliffs and talus above streams | |||||||||
Elevation | 700–1400 m (2300–4600 ft) | |||||||||
Distribution |
OR
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CA; ID; MT; NV; OR; UT; WA; WY; AB; BC
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Discussion | Drymocallis campanulata is one of the more distinct species in the genus, with its campanulate flowers, relatively large butter yellow petals, and densely glandular pedicels. It occurs primarily in the canyons of the John Day River in Grant and Wheeler counties. The invalidly published Potentilla campanulata D. D. Keck provides a full description and illustration for the subsequent basionym. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 3 (3 in the flora). Drymocallis pseudorupestris occurs from Alberta and Washington to California and Utah, mostly in montane habitats; it is the species most often associated with rocky habitats, including talus slopes, for which its relatively elongate caudex branches are an obvious adaptation. Vestiture is dominated by abundant septate glands on stems and in the inflorescences. Except for var. pseudorupestris, which occurs only in the northeastern part of the species range, plants are relatively short, usually less than 2.5 dm. Three intergrading varieties accommodate the extremes at the northeastern and southern ends of the range. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 294. | FNA vol. 9, p. 289. | ||||||||
Parent taxa | ||||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Potentilla glandulosa var. campanulata | Potentilla pseudorupestris, D. glandulosa subsp. pseudorupestris, P. glandulosa subsp. pseudorupestris, P. glandulosa var. pseudorupestris, P. rupestris var. americana | ||||||||
Name authority | (C. L. Hitchcock) Ertter: J. Bot. Res. Inst. Texas 1: 43. (2007) | (Rydberg) Rydberg: Monogr. N. Amer. Potentilleae, 194. (1898) | ||||||||
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