Drymocallis campanulata |
Drymocallis convallaria |
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John Day drymocallis, John Day drymocallis or wood beauty |
cordilleran drymocallis or wood beauty, cordilleran woodbeauty, cream cinquefoil, sharp tooth cinquefoil, valley cinquefoil, white cinquefoil |
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Caudex branches | elongate. |
short. |
Stems | openly tufted to loosely spaced, 1.5–4.5 dm; base 1.2–3.5 mm diam., ± densely septate-glandular. |
usually solitary, sometimes tufted, 4–9(–10) dm; base 2–5 mm diam., ± densely septate-glandular. |
Leaves | sparsely to ± densely hairy; basal 6–20 cm, leaflet pairs (2–)3–4(–5); terminal leaflet broadly obovate to nearly round, 1–4 × 1–3.5 cm, teeth usually ± double, 5–10 per side, apex rounded; cauline 0–2, well developed, leaflet pairs 2–3. |
sparsely to moderately, rarely densely, hairy; basal (10–)15–25(–30) cm, leaflet pairs (2–)3–4; terminal leaflet usually obovate, sometimes elliptic, 3–5(–6) × 1.5–4.5 cm, teeth double, 9–16 per side, apex rounded-obtuse; cauline (1–)2–4, well developed at least proximally, leaflet pairs usually 3. |
Inflorescences | 5–50-flowered, leafy, open, (1/5–)1/2(–4/5) of stem, wide, branch angles 25–50°. |
(5–)10–40-flowered, not leafy, congested to open, (1/8–)1/6–1/3(–1/2) of stem, narrow, branch angles 5–20°. |
Pedicels | 2–5 (proximal to 10) mm, sparsely to moderately short-hairy, predominantly septate-glandular. |
1–5 (proximal to 25) mm, sparsely to ± densely short-hairy, predominantly septate-glandular. |
Flowers | opening narrowly; epicalyx bractlets usually lanceolate, sometimes linear, 3–4 × 1–1.5 mm; sepals ± erect, 5–8 mm, apex acute to obtuse; petals overlapping, ± erect, light yellow, broadly obovate, 5–11 × (3–)5–8 mm, equal to or exceeding sepals; filaments 1.5–3 mm, anthers 1 mm; styles slender, (1–)1.5–2.5 mm. |
opening ± widely; epicalyx bractlets narrowly elliptic, (2–)4–6 × 0.7–1.5 mm; sepals ± spreading, (4–)5–8(–10) mm, apex acute; petals not or scarcely overlapping, ± spreading, cream-white to pale yellow, obovate-elliptic, sometimes broadly so, (3–)5–8 × (2–)4–6 mm, ± equal to sepals; filaments 1.5–2.5 mm, anthers 0.7–1 mm; styles thickened, 0.8 mm. |
Achenes | light brown, 1 mm. |
light brown, 1 mm. |
Drymocallis campanulata |
Drymocallis convallaria |
|
Phenology | Flowering May–Jul. | Flowering (May–)Jun–Aug. |
Habitat | Basaltic cliffs and talus above streams | Dry to seasonally moist meadows, open forests, sagebrush and grassy rocky slopes |
Elevation | 700–1400 m (2300–4600 ft) | 100–3000 m (300–9800 ft) |
Distribution |
OR
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AK; AZ; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC; YT
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Discussion | Drymocallis campanulata is one of the more distinct species in the genus, with its campanulate flowers, relatively large butter yellow petals, and densely glandular pedicels. It occurs primarily in the canyons of the John Day River in Grant and Wheeler counties. The invalidly published Potentilla campanulata D. D. Keck provides a full description and illustration for the subsequent basionym. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Drymocallis convallaria is widespread in western North America, occurring mostly west of the Continental Divide from south-central Alaska to Arizona. It differs from most sympatric species in usually having a single thick stem, a narrow inflorescence, and cream-white to pale yellow petals, at most only slightly longer than the sepals. Some collections from southern Idaho and northern Nevada that have very small petals in the size range of Drymocallis micropetala are included here on the basis of glandular pedicels. Excluded here are comparably small-petaled plants from the Warner Mountains of California and Oregon that have the aspect of D. convallaria but the blunter, redder achenes of D. glandulosa. Populations in northern Idaho and adjacent Washington that approach D. arguta in size and vestiture are provisionally treated as D. convallaria on the basis of leaflet shape. Specimens from New Mexico previously assigned to this taxon have largely been redetermined to D. arguta in the strict sense. See also discussions of Potentilla fissa var. major under 1. D. fissa, and of D. valida under the genus discussion. The illegitimate names P. glutinosa Nuttall ex Rydberg and D. glutinosa Rydberg have been applied most often to D. convallaria. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 294. | FNA vol. 9, p. 286. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Potentilla glandulosa var. campanulata | Potentilla convallaria, D. arguta subsp. convallaria, P. arguta subsp. convallaria, P. arguta var. convallaria |
Name authority | (C. L. Hitchcock) Ertter: J. Bot. Res. Inst. Texas 1: 43. (2007) | (Rydberg) Rydberg: Monogr. N. Amer. Potentilleae, 193. (1898) |
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