Drymocallis ashlandica |
Rosaceae tribe Potentilleae |
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Ashland cinquefoil, Mazama drymocallis or wood beauty |
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Habit | Herbs, perennial, rarely annual or biennial, shrubs, or subshrubs; unarmed. | |||||||||||||||||||||||||||||||||||||||||||||||||||||
Caudex branches | short. |
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Stems | tufted, (1–)2–5 dm; base 2–3 mm diam., ± densely septate-glandular. |
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Leaves | moderately to ± densely hairy; basal 6–16 cm, leaflet pairs 2–3(–4); terminal leaflet broadly obovate, 1.5–4 × 1–3 cm, teeth ± double, 7–12 per side, apex rounded; cauline 0–2, moderately developed, leaflet pairs 2–3. |
alternate, rarely opposite, pinnately (palmately) compound (simple in Alchemilla, Aphanes, and Chamaerhodos); stipules persistent (absent in Chamaerhodos), adnate to petiole; venation pinnate or palmate. |
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Inflorescences | 5–15(–20)-flowered, not leafy, congested to deeply branched, 1/10–2/3 or less of stem, usually narrow, rarely ± wide, branch angles (5–)10–20(–30)°. |
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Pedicels | 2–10 (proximal to 30) mm, sparsely to moderately short-hairy, predominantly septate-glandular, often obscuring other vestiture. |
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Flowers | opening widely; epicalyx bractlets linear to broadly elliptic, 3–5 × 1–2 mm; sepals spreading, 5–7(–9) mm, apex acute; petals overlapping, spreading, light yellow, broadly obovate, 5–9 × 4–7 mm, equal to or longer than sepals; filaments 1.5–5 mm, anthers 0.7–1.2 mm; styles ± thickened, 1 mm. |
perianth and androecium perigynous; epicalyx bractlets present, sometimes absent; hypanthium usually patelliform, cupulate, or campanulate, sometimes turbinate, saucer-shaped, flat-bottomed, or subglobose to ellipsoid or ovoid; torus flat to conic or turbinate, enlarged (absent or reduced in Alchemilla, Aphanes, and Chamaerhodos); carpels 1–260, styles basal or lateral to subterminal, distinct; ovules 1(or 2), basal. |
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Fruits | aggregated achenes (achenes in Alchemilla and Aphanes); torus sometimes fleshy; styles deciduous or persistent, not elongate. |
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Achenes | light brown, 1 mm. |
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Drymocallis ashlandica |
Rosaceae tribe Potentilleae |
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Phenology | Flowering Jun–Aug. | |||||||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Meadows, roadsides, sandy shorelines | |||||||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 500–2000 m (1600–6600 ft) | |||||||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
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North America; Mexico; Central America; South America; West Indies; Bermuda; Eurasia; Africa; Atlantic Islands; Pacific Islands; Australia |
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Discussion | The name Potentilla glandulosa subsp. ashlandica was misapplied to Drymocallis lactea var. austiniae (B. Ertter 1993); as circumscribed here, D. ashlandica is a poorly known species restricted to the Cascade and Siskiyou ranges of southwestern Oregon. It very likely also occurs in adjacent California; no collections from there are known. Drymocallis ashlandica differs from most sympatric populations of D. lactea var. austiniae and resembles D. pseudorupestris in its densely glandular inflorescences, which tend to be more densely congested and/or narrower than those of the other two species. Drymocallis ashlandica was described by Greene as Potentilla ashlandica, and by Howell as P. ciliata (a later homonym), both based on a collection by Howell from the Siskiyou Mountains near Ashland, Oregon. The circumscription here, which matches that of D. D. Keck (in J. Clausen et al. 1940) and M. E. Peck (1961), encompasses populations from near Oregon Caves and Ashland in the Siskiyou Mountains to near McKenzie Bridge, Lane County, and Sparks Lake, Deschutes County. It is currently known from relatively few collections, mostly from the Crater Lake area, and may prove to be of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 14–22, species ca. 860 (14 genera, 189 species, including 1 hybrid, in the flora area). The base chromosome number for Potentilleae is mostly x = 7 (8 in Alchemilla and Aphanes; 14 in Comarum). Variation in the number of genera recognized in Potentilleae is due to differences in generic delimitation between D. Potter et al. (2007) and the authors of Potentilla and segregates here (see 9. Ivesia and 8. Potentilla for discussion). In the former, Duchesnea, Horkelia, Horkeliella, and Ivesia are included within Potentilla. Likewise, Aphanes is included within Alchemilla by Potter et al. while it is kept distinct here. Potentilla and its segregates and Fragaria are host to Phragmidium rusts, but not the other genera of the tribe. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 288. | FNA vol. 9, p. 119. | ||||||||||||||||||||||||||||||||||||||||||||||||||||
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis | Rosaceae > subfam. Rosoideae | ||||||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | Potentilla ashlandica, D. glandulosa subsp. ashlandica, P. glandulosa subsp. ashlandica | |||||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | (Greene) Rydberg: Monogr. N. Amer. Potentilleae, 200. (1898) | Sweet: Brit. Fl. Gard. 2: sub plate 124. (1825) | ||||||||||||||||||||||||||||||||||||||||||||||||||||
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