Drymocallis ashlandica |
Drymocallis glandulosa |
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Ashland cinquefoil, Mazama drymocallis or wood beauty |
Douglas' woodbeauty, Greene's drymocallis, sticky cinquefoil |
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Caudex branches | short. |
short. |
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Stems | tufted, (1–)2–5 dm; base 2–3 mm diam., ± densely septate-glandular. |
tufted, (1–)1.5–7.5 dm; base 1–5 mm diam., moderately to densely septate-glandular. |
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Leaves | moderately to ± densely hairy; basal 6–16 cm, leaflet pairs 2–3(–4); terminal leaflet broadly obovate, 1.5–4 × 1–3 cm, teeth ± double, 7–12 per side, apex rounded; cauline 0–2, moderately developed, leaflet pairs 2–3. |
sparsely to moderately hairy; basal (3–)6–30 cm, leaflet pairs (1–)2–4; terminal leaflet broadly obovate to rhombic-elliptic, (1–)1.5–7(–8) × 1–5 cm, teeth usually double (irregularly single in var. viscida), (4–)6–18(–20) per side, apex rounded to obtuse; cauline 0–3, developed or reduced, leaflet pairs 1–4. |
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Inflorescences | 5–15(–20)-flowered, not leafy, congested to deeply branched, 1/10–2/3 or less of stem, usually narrow, rarely ± wide, branch angles (5–)10–20(–30)°. |
(2–)5–40-flowered, leafy or not, open or of congested clusters terminating branches, (1/10–)1/5–3/4 of stem, usually wide (narrow in var. viscida), branch angles (15–)20–55°. |
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Pedicels | 2–10 (proximal to 30) mm, sparsely to moderately short-hairy, predominantly septate-glandular, often obscuring other vestiture. |
1–10 (proximal to 30) mm, not or sparsely short-hairy, predominantly septate-glandular. |
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Flowers | opening widely; epicalyx bractlets linear to broadly elliptic, 3–5 × 1–2 mm; sepals spreading, 5–7(–9) mm, apex acute; petals overlapping, spreading, light yellow, broadly obovate, 5–9 × 4–7 mm, equal to or longer than sepals; filaments 1.5–5 mm, anthers 0.7–1.2 mm; styles ± thickened, 1 mm. |
opening widely; epicalyx bractlets linear-lanceolate to elliptic, 1.5–7(–8) × 0.5–2.5(–3) mm; sepals spreading or reflexed, (3–)4–11 mm, apex usually broadly obtuse with mucronate tip, sometimes acute; petals not overlapping, spreading or reflexed, cream-white to yellow, narrowly to broadly obovate or ovate to nearly round, 2–6.5 × 1.5–5.5 mm, shorter than or equal to sepals; filaments 0.5–2 mm, anthers 0.5–1 mm; styles thickened, 1–1.5 mm. |
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Achenes | light brown, 1 mm. |
reddish brown, 0.9–1.5 mm. |
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Drymocallis ashlandica |
Drymocallis glandulosa |
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Phenology | Flowering Jun–Aug. | |||||||||||||
Habitat | Meadows, roadsides, sandy shorelines | |||||||||||||
Elevation | 500–2000 m (1600–6600 ft) | |||||||||||||
Distribution |
OR
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AZ; CA; ID; MT; NV; OR; UT; WA; BC; nw Mexico
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Discussion | The name Potentilla glandulosa subsp. ashlandica was misapplied to Drymocallis lactea var. austiniae (B. Ertter 1993); as circumscribed here, D. ashlandica is a poorly known species restricted to the Cascade and Siskiyou ranges of southwestern Oregon. It very likely also occurs in adjacent California; no collections from there are known. Drymocallis ashlandica differs from most sympatric populations of D. lactea var. austiniae and resembles D. pseudorupestris in its densely glandular inflorescences, which tend to be more densely congested and/or narrower than those of the other two species. Drymocallis ashlandica was described by Greene as Potentilla ashlandica, and by Howell as P. ciliata (a later homonym), both based on a collection by Howell from the Siskiyou Mountains near Ashland, Oregon. The circumscription here, which matches that of D. D. Keck (in J. Clausen et al. 1940) and M. E. Peck (1961), encompasses populations from near Oregon Caves and Ashland in the Siskiyou Mountains to near McKenzie Bridge, Lane County, and Sparks Lake, Deschutes County. It is currently known from relatively few collections, mostly from the Crater Lake area, and may prove to be of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 4 (4 in the flora). Even with the removal of most of the subspecies of Potentilla glandulosa treated by Keck, Drymocallis glandulosa encompasses an inordinate range of variation. This is because of the apparently clinal variation between the coastal extreme (var. wrangelliana), with leafy inflorescences and broad cream-white petals, the montane var. reflexa, with less leafy inflorescences and narrow yellow petals, and the transitional interior var. glandulosa. Variety reflexa, in turn, intergrades in the mountains of southern California with var. viscida, which is anomalous in D. glandulosa with its narrow inflorescences and irregularly single-toothed leaflets. Variation outside of California is unclear, with no evident pattern to differences in petal size and color; these have largely been assigned to var. glandulosa. At its most distinctive, Drymocallis glandulosa is a robust, densely glandular plant with widely branched leafy inflorescences (except var. viscida), relatively short, stout pedicels, broad sepals with rounded, mucronate tips (except var. viscida and sometimes var. reflexa) that are often significantly enlarged in fruit, and non-overlapping petals that are shorter than to scarcely equal to the sepals. The relatively large achenes are blunter and redder than those of most other species of Drymocallis, as discussed by J. Clausen et al. (1940). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 288. | FNA vol. 9, p. 292. | ||||||||||||
Parent taxa | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis | Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis | ||||||||||||
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Synonyms | Potentilla ashlandica, D. glandulosa subsp. ashlandica, P. glandulosa subsp. ashlandica | Potentilla glandulosa | ||||||||||||
Name authority | (Greene) Rydberg: Monogr. N. Amer. Potentilleae, 200. (1898) | (Lindley) Rydberg: Monogr. N. Amer. Potentilleae, 198. (1898) | ||||||||||||
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