Flora of North America species comparison

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Drymocallis ashlandica

Drymocallis cuneifolia

Ashland cinquefoil, Mazama drymocallis or wood beauty
Caudex branches

short.

elongate.
Stems

tufted, (1–)2–5 dm;

base 2–3 mm diam., ± densely septate-glandular.

openly tufted to loosely spaced, 0.5–4.5 dm;

base 1–2 mm diam., sparsely septate-glandular.
Leaves

moderately to ± densely hairy;

basal 6–16 cm, leaflet pairs 2–3(–4);

terminal leaflet broadly obovate, 1.5–4 × 1–3 cm, teeth ± double, 7–12 per side, apex rounded;

cauline 0–2, moderately developed, leaflet pairs 2–3.

sparsely to moderately hairy;

basal 2–15 cm, leaflet pairs (2–)3–5;

terminal leaflet broadly cuneate to nearly round, 0.6–2(–3.5) × 0.5–2 cm, teeth irregularly single, 2–4 per side, apex rounded to truncate;

cauline 0–1(–2), reduced, leaflet pairs 1–2.
Inflorescences

5–15(–20)-flowered, not leafy, congested to deeply branched, 1/10–2/3 or less of stem, usually narrow, rarely ± wide, branch angles (5–)10–20(–30)°.

3–10(–15)-flowered, not leafy, open, 1/4–3/4 of stem, wide, branch angles 20–75°.
Pedicels

2–10 (proximal to 30) mm, sparsely to moderately short-hairy, predominantly septate-glandular, often obscuring other vestiture.

2–15 (proximal to 30) mm, sparsely to moderately short-hairy, septate-glandular.
Flowers

opening widely;

epicalyx bractlets linear to broadly elliptic, 3–5 × 1–2 mm;

sepals spreading, 5–7(–9) mm, apex acute;

petals overlapping, spreading, light yellow, broadly obovate, 5–9 × 4–7 mm, equal to or longer than sepals;

filaments 1.5–5 mm, anthers 0.7–1.2 mm;

styles ± thickened, 1 mm.

opening narrowly;

epicalyx bractlets linear to narrowly elliptic or lanceolate, 1–2 × 0.5 mm;

sepals erect, 2–5 mm, apex broadly obtuse;

petals scarcely overlapping, erect, yellow, narrowly obovate, 2–4 × 1.5–2.5 mm, ± equal to or slightly longer than sepals;

filaments 1.5–2.5 mm, anthers 0.5–0.8 mm;

styles slender, 1.5–2.5 mm.
Achenes

light brown, 1 mm.

brown, 0.8–1.2 mm.

Drymocallis ashlandica

Drymocallis cuneifolia
Phenology Flowering Jun–Aug.
Habitat Meadows, roadsides, sandy shorelines
Elevation 500–2000 m (1600–6600 ft)
Distribution
from FNA
OR
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

The name Potentilla glandulosa subsp. ashlandica was misapplied to Drymocallis lactea var. austiniae (B. Ertter 1993); as circumscribed here, D. ashlandica is a poorly known species restricted to the Cascade and Siskiyou ranges of southwestern Oregon. It very likely also occurs in adjacent California; no collections from there are known. Drymocallis ashlandica differs from most sympatric populations of D. lactea var. austiniae and resembles D. pseudorupestris in its densely glandular inflorescences, which tend to be more densely congested and/or narrower than those of the other two species.

Drymocallis ashlandica was described by Greene as Potentilla ashlandica, and by Howell as P. ciliata (a later homonym), both based on a collection by Howell from the Siskiyou Mountains near Ashland, Oregon. The circumscription here, which matches that of D. D. Keck (in J. Clausen et al. 1940) and M. E. Peck (1961), encompasses populations from near Oregon Caves and Ashland in the Siskiyou Mountains to near McKenzie Bridge, Lane County, and Sparks Lake, Deschutes County. It is currently known from relatively few collections, mostly from the Crater Lake area, and may prove to be of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Drymocallis cuneifolia has a complex nomenclatural history, complicated by the extreme rarity of the typical variety. Rydberg described the species on the basis of a single collection from the San Bernardino Mountains of San Bernardino County, probably near Green Lead Mines. When comparable (though significantly smaller) plants were found in the San Gabriel Mountains of Los Angeles County, P. A. Munz and I. M. Johnston (1925) adopted Potentilla cuneifolia (Rydberg) Th. Wolf for both extremes; this name was later replaced with P. peirsonii because of the earlier P. cuneifolia Bertoloni. In the absence of comparable new collections from the San Bernardino Mountains, D. D. Keck (in J. Clausen et al. 1940) concluded that the type of D. cuneifolia was merely an immature specimen of D. lactea var. lactea and accordingly described P. glandulosa subsp. ewanii to accommodate populations in the San Gabriel Mountains.

The 2004 discovery of a small population of plants comparable to the type of Drymocallis cuneifolia from near the type locality (Elvin 3555, IRVC, UCR) confirms that D. cuneifolia is a valid taxon that shares most of the diagnostic features of populations in the San Gabriel Mountains. Plants in the San Bernardino Mountains tend to be larger than those in the San Gabriel Mountains, with fewer and somewhat differently shaped leaflets; they are accordingly treated here as varieties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Key
1. Stems 2–4.5 dm; basal leaves 5–15 cm; San Bernardino Mountains.
var. cuneifolia
1. Stems 0.5–2(–2.5) dm; basal leaves 2–10 cm; San Gabriel Mountains.
var. ewanii
Source FNA vol. 9, p. 288. FNA vol. 9, p. 294.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis
Sibling taxa
D. arguta, D. arizonica, D. campanulata, D. convallaria, D. cuneifolia, D. deseretica, D. fissa, D. glabrata, D. glandulosa, D. hansenii, D. lactea, D. micropetala, D. pseudorupestris, D. rhomboidea
D. arguta, D. arizonica, D. ashlandica, D. campanulata, D. convallaria, D. deseretica, D. fissa, D. glabrata, D. glandulosa, D. hansenii, D. lactea, D. micropetala, D. pseudorupestris, D. rhomboidea
Subordinate taxa
D. cuneifolia var. cuneifolia, D. cuneifolia var. ewanii
Synonyms Potentilla ashlandica, D. glandulosa subsp. ashlandica, P. glandulosa subsp. ashlandica Potentilla peirsonii
Name authority (Greene) Rydberg: Monogr. N. Amer. Potentilleae, 200. (1898) Rydberg: Monogr. N. Amer. Potentilleae, 204, plate 111 [as cuneata]. (1898)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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