FNA: Drymocallis ashlandica vs. Drymocallis arizonica

	Drymocallis ashlandica	Drymocallis arizonica
	Ashland cinquefoil, Mazama drymocallis or wood beauty	Arizona drymocallis or wood beauty
Caudex branches	short.	short.
Stems	tufted, (1–)2–5 dm; base 2–3 mm diam., ± densely septate-glandular.	± tufted, (1.5–)2.5–6 dm; base 1.5–2.5 mm diam., sparsely to densely septate-glandular.
Leaves	moderately to ± densely hairy; basal 6–16 cm, leaflet pairs 2–3(–4); terminal leaflet broadly obovate, 1.5–4 × 1–3 cm, teeth ± double, 7–12 per side, apex rounded; cauline 0–2, moderately developed, leaflet pairs 2–3.	sparsely to moderately hairy; basal 5–22 cm, leaflet pairs (2–)3–4(–5); terminal leaflet broadly obovate-elliptic, 1–5 × 1–2.5(–4) cm, teeth double, 6–18 per side, apex rounded to acute; cauline 1–4, well developed, leaflet pairs 2–3.
Inflorescences	5–15(–20)-flowered, not leafy, congested to deeply branched, 1/10–2/3 or less of stem, usually narrow, rarely ± wide, branch angles (5–)10–20(–30)°.	5–30-flowered, not notably leafy, compact, 1/10–1/4(–1/2) of stem, narrow, branch angles 5–25°.
Pedicels	2–10 (proximal to 30) mm, sparsely to moderately short-hairy, predominantly septate-glandular, often obscuring other vestiture.	1–10 (proximal to 20) mm, sparsely to ± densely short-hairy, predominantly septate-glandular.
Flowers	opening widely; epicalyx bractlets linear to broadly elliptic, 3–5 × 1–2 mm; sepals spreading, 5–7(–9) mm, apex acute; petals overlapping, spreading, light yellow, broadly obovate, 5–9 × 4–7 mm, equal to or longer than sepals; filaments 1.5–5 mm, anthers 0.7–1.2 mm; styles ± thickened, 1 mm.	opening widely; epicalyx bractlets linear-lanceolate to narrowly elliptic, 2–4(–7) × 1–2 mm; sepals spreading, 4–6(–11) mm, apex ± acute; petals not or scarcely overlapping, spreading, cream-white, ± obovate, 3–6 × 2.5–5 mm, shorter than or equal to sepals; filaments 1–3 mm, anthers 0.7–1 mm; styles thickened, 1 mm.
Achenes	light brown, 1 mm.	light brown, 1 mm.
2n		= 14.

	Drymocallis ashlandica	Drymocallis arizonica
Phenology	Flowering Jun–Aug.	Flowering May–Jul.
Habitat	Meadows, roadsides, sandy shorelines	Streamsides, rocky sites, open forest floors, pine and aspen forests
Elevation	500–2000 m (1600–6600 ft)	1800–3200 m (5900–10500 ft)
Distribution	OR [WildflowerSearch map] [BONAP county map]	AZ; UT [BONAP county map]
Discussion	The name Potentilla glandulosa subsp. ashlandica was misapplied to Drymocallis lactea var. austiniae (B. Ertter 1993); as circumscribed here, D. ashlandica is a poorly known species restricted to the Cascade and Siskiyou ranges of southwestern Oregon. It very likely also occurs in adjacent California; no collections from there are known. Drymocallis ashlandica differs from most sympatric populations of D. lactea var. austiniae and resembles D. pseudorupestris in its densely glandular inflorescences, which tend to be more densely congested and/or narrower than those of the other two species. Drymocallis ashlandica was described by Greene as Potentilla ashlandica, and by Howell as P. ciliata (a later homonym), both based on a collection by Howell from the Siskiyou Mountains near Ashland, Oregon. The circumscription here, which matches that of D. D. Keck (in J. Clausen et al. 1940) and M. E. Peck (1961), encompasses populations from near Oregon Caves and Ashland in the Siskiyou Mountains to near McKenzie Bridge, Lane County, and Sparks Lake, Deschutes County. It is currently known from relatively few collections, mostly from the Crater Lake area, and may prove to be of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Drymocallis arizonica encompasses populations in Arizona north of the Mogollon Rim and in Utah as far north as Garfield and Sevier counties, including the Henry Mountains. It is most distinctive in northern Arizona, where plants commonly have basal leaves with four pairs of lateral leaflets and compact inflorescences. The species intergrades with D. convallaria but is in general shorter and more likely multistemmed. It also shares similarities with D. pseudorupestris but has more developed cauline leaves and a more compact inflorescence. The intergrade zone between all three species and D. deseretica in southwestern Utah is particularly problematic and perhaps indicative of yet additional taxa deserving recognition. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 9, p. 288.	FNA vol. 9, p. 284.
Parent taxa	Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis	Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis
Sibling taxa	D. arguta, D. arizonica, D. campanulata, D. convallaria, D. cuneifolia, D. deseretica, D. fissa, D. glabrata, D. glandulosa, D. hansenii, D. lactea, D. micropetala, D. pseudorupestris, D. rhomboidea	D. arguta, D. ashlandica, D. campanulata, D. convallaria, D. cuneifolia, D. deseretica, D. fissa, D. glabrata, D. glandulosa, D. hansenii, D. lactea, D. micropetala, D. pseudorupestris, D. rhomboidea
Synonyms	Potentilla ashlandica, D. glandulosa subsp. ashlandica, P. glandulosa subsp. ashlandica	D. glandulosa subsp. arizonica, Potentilla glandulosa subsp. arizonica, P. macdougalii
Name authority	(Greene) Rydberg: Monogr. N. Amer. Potentilleae, 200. (1898)	Rydberg: in N. L. Britton et al., N. Amer. Fl. 22: 373. (1908)
Web links	Local floras: OR Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Drymocallis arizonica

Ashland cinquefoil, Mazama drymocallis or wood beauty

Arizona drymocallis or wood beauty

Caudex branches

short.

Stems

tufted, (1–)2–5 dm;

base 2–3 mm diam., ± densely septate-glandular.

± tufted, (1.5–)2.5–6 dm;

base 1.5–2.5 mm diam., sparsely to densely septate-glandular.

Leaves

moderately to ± densely hairy;

basal 6–16 cm, leaflet pairs 2–3(–4);

terminal leaflet broadly obovate, 1.5–4 × 1–3 cm, teeth ± double, 7–12 per side, apex rounded;

cauline 0–2, moderately developed, leaflet pairs 2–3.

sparsely to moderately hairy;

basal 5–22 cm, leaflet pairs (2–)3–4(–5);

terminal leaflet broadly obovate-elliptic, 1–5 × 1–2.5(–4) cm, teeth double, 6–18 per side, apex rounded to acute;

cauline 1–4, well developed, leaflet pairs 2–3.

Inflorescences

5–15(–20)-flowered, not leafy, congested to deeply branched, 1/10–2/3 or less of stem, usually narrow, rarely ± wide, branch angles (5–)10–20(–30)°.

5–30-flowered, not notably leafy, compact, 1/10–1/4(–1/2) of stem, narrow, branch angles 5–25°.

Pedicels

2–10 (proximal to 30) mm, sparsely to moderately short-hairy, predominantly septate-glandular, often obscuring other vestiture.

1–10 (proximal to 20) mm, sparsely to ± densely short-hairy, predominantly septate-glandular.

Flowers

opening widely;

epicalyx bractlets linear to broadly elliptic, 3–5 × 1–2 mm;

sepals spreading, 5–7(–9) mm, apex acute;

petals overlapping, spreading, light yellow, broadly obovate, 5–9 × 4–7 mm, equal to or longer than sepals;

filaments 1.5–5 mm, anthers 0.7–1.2 mm;

styles ± thickened, 1 mm.

opening widely;

epicalyx bractlets linear-lanceolate to narrowly elliptic, 2–4(–7) × 1–2 mm;

sepals spreading, 4–6(–11) mm, apex ± acute;

petals not or scarcely overlapping, spreading, cream-white, ± obovate, 3–6 × 2.5–5 mm, shorter than or equal to sepals;

filaments 1–3 mm, anthers 0.7–1 mm;

styles thickened, 1 mm.

Achenes

light brown, 1 mm.

= 14.

Drymocallis arizonica

Phenology

Flowering Jun–Aug.

Flowering May–Jul.

Habitat

Meadows, roadsides, sandy shorelines

Streamsides, rocky sites, open forest floors, pine and aspen forests

Elevation

500–2000 m (1600–6600 ft)

1800–3200 m (5900–10500 ft)

Distribution

[WildflowerSearch map]

[BONAP county map]

AZ; UT

[BONAP county map]

Discussion

The name Potentilla glandulosa subsp. ashlandica was misapplied to Drymocallis lactea var. austiniae (B. Ertter 1993); as circumscribed here, D. ashlandica is a poorly known species restricted to the Cascade and Siskiyou ranges of southwestern Oregon. It very likely also occurs in adjacent California; no collections from there are known. Drymocallis ashlandica differs from most sympatric populations of D. lactea var. austiniae and resembles D. pseudorupestris in its densely glandular inflorescences, which tend to be more densely congested and/or narrower than those of the other two species.

Drymocallis ashlandica was described by Greene as Potentilla ashlandica, and by Howell as P. ciliata (a later homonym), both based on a collection by Howell from the Siskiyou Mountains near Ashland, Oregon. The circumscription here, which matches that of D. D. Keck (in J. Clausen et al. 1940) and M. E. Peck (1961), encompasses populations from near Oregon Caves and Ashland in the Siskiyou Mountains to near McKenzie Bridge, Lane County, and Sparks Lake, Deschutes County. It is currently known from relatively few collections, mostly from the Crater Lake area, and may prove to be of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Drymocallis arizonica encompasses populations in Arizona north of the Mogollon Rim and in Utah as far north as Garfield and Sevier counties, including the Henry Mountains. It is most distinctive in northern Arizona, where plants commonly have basal leaves with four pairs of lateral leaflets and compact inflorescences. The species intergrades with D. convallaria but is in general shorter and more likely multistemmed. It also shares similarities with D. pseudorupestris but has more developed cauline leaves and a more compact inflorescence. The intergrade zone between all three species and D. deseretica in southwestern Utah is particularly problematic and perhaps indicative of yet additional taxa deserving recognition.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 9, p. 288.

FNA vol. 9, p. 284.

Parent taxa

Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis

Sibling taxa

D. arguta, D. arizonica, D. campanulata, D. convallaria, D. cuneifolia, D. deseretica, D. fissa, D. glabrata, D. glandulosa, D. hansenii, D. lactea, D. micropetala, D. pseudorupestris, D. rhomboidea

D. arguta, D. ashlandica, D. campanulata, D. convallaria, D. cuneifolia, D. deseretica, D. fissa, D. glabrata, D. glandulosa, D. hansenii, D. lactea, D. micropetala, D. pseudorupestris, D. rhomboidea

Synonyms

Potentilla ashlandica, D. glandulosa subsp. ashlandica, P. glandulosa subsp. ashlandica

D. glandulosa subsp. arizonica, Potentilla glandulosa subsp. arizonica, P. macdougalii

Name authority

(Greene) Rydberg: Monogr. N. Amer. Potentilleae, 200. (1898)

Rydberg: in N. L. Britton et al., N. Amer. Fl. 22: 373. (1908)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora

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Drymocallis ashlandica

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Drymocallis ashlandica

Drymocallis arizonica