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Arizona drymocallis or wood beauty

Habit Herbs, shrubs, or subshrubs.
Caudex branches

short.

Stems

± tufted, (1.5–)2.5–6 dm;

base 1.5–2.5 mm diam., sparsely to densely septate-glandular.

Leaves

sparsely to moderately hairy;

basal 5–22 cm, leaflet pairs (2–)3–4(–5);

terminal leaflet broadly obovate-elliptic, 1–5 × 1–2.5(–4) cm, teeth double, 6–18 per side, apex rounded to acute;

cauline 1–4, well developed, leaflet pairs 2–3.

alternate, rarely opposite, pinnately compound, sometimes simple or palmately compound;

stipules present, rarely absent.

Inflorescences

5–30-flowered, not notably leafy, compact, 1/10–1/4(–1/2) of stem, narrow, branch angles 5–25°.

Pedicels

1–10 (proximal to 20) mm, sparsely to ± densely short-hairy, predominantly septate-glandular.

Flowers

opening widely;

epicalyx bractlets linear-lanceolate to narrowly elliptic, 2–4(–7) × 1–2 mm;

sepals spreading, 4–6(–11) mm, apex ± acute;

petals not or scarcely overlapping, spreading, cream-white, ± obovate, 3–6 × 2.5–5 mm, shorter than or equal to sepals;

filaments 1–3 mm, anthers 0.7–1 mm;

styles thickened, 1 mm.

torus usually enlarged, sometimes small or absent;

carpels 1–260(–450), distinct, free, styles distinct, rarely connate (Roseae);

ovules 1(or 2), collateral (Rubeae) or superposed (Fallugia, Filipendula).

Fruits

achenes or aggregated achenes sometimes with fleshy, urn-shaped hypanthium or enlarged torus, sometimes aggregated drupelets;

styles persistent or deciduous, not elongate (elongate but not plumose in Geum).

Achenes

light brown, 1 mm.

x

= 7(8).

2n

= 14.

Drymocallis arizonica

Rosaceae subfam. rosoideae

Phenology Flowering May–Jul.
Habitat Streamsides, rocky sites, open forest floors, pine and aspen forests
Elevation 1800–3200 m (5900–10500 ft)
Distribution
from FNA
AZ; UT
[BONAP county map]
North America; Mexico; Central America; South America; West Indies; Bermuda; Eurasia; Africa; Atlantic Islands; Indian Ocean Islands; Pacific Islands; Australia
Discussion

Drymocallis arizonica encompasses populations in Arizona north of the Mogollon Rim and in Utah as far north as Garfield and Sevier counties, including the Henry Mountains. It is most distinctive in northern Arizona, where plants commonly have basal leaves with four pairs of lateral leaflets and compact inflorescences. The species intergrades with D. convallaria but is in general shorter and more likely multistemmed. It also shares similarities with D. pseudorupestris but has more developed cauline leaves and a more compact inflorescence. The intergrade zone between all three species and D. deseretica in southwestern Utah is particularly problematic and perhaps indicative of yet additional taxa deserving recognition.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Variation in the number of genera in subfam. Rosoideae is due to differences in generic delimitation between D. Potter et al. (2007) and the authors of some Potentilleae genera. Cyanogenic glycosides and sorbitol are absent in the subfamily.

Tribes 6, genera 28–35, species ca. 1600 (6 tribes, 26 genera, 302 species, including 1 hybrid, in the flora)

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 284. FNA vol. 9, p. 23. Author: Luc Brouillet.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Potentilleae > Drymocallis Rosaceae
Sibling taxa
D. arguta, D. ashlandica, D. campanulata, D. convallaria, D. cuneifolia, D. deseretica, D. fissa, D. glabrata, D. glandulosa, D. hansenii, D. lactea, D. micropetala, D. pseudorupestris, D. rhomboidea
Subordinate taxa
Synonyms D. glandulosa subsp. arizonica, Potentilla glandulosa subsp. arizonica, P. macdougalii
Name authority Rydberg: in N. L. Britton et al., N. Amer. Fl. 22: 373. (1908) Arnott: Botany, 107. (1832)
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